V. Grabovsky, S. Feigelson, C. Chen, D. Bleijs, and A. Peled, Subsecond Induction of ??4 Integrin Clustering by Immobilized Chemokines Stimulates Leukocyte Tethering and Rolling on Endothelial Vascular Cell Adhesion Molecule 1 under Flow Conditions, The Journal of Experimental Medicine, vol.161, issue.4, pp.495-506, 2000.
DOI : 10.1161/01.RES.82.1.30

R. Mcever and R. Cummings, Perspectives series: cell adhesion in vascular biology. Role of PSGL-1 binding to selectins in leukocyte recruitment., Journal of Clinical Investigation, vol.100, issue.3, pp.485-491, 1997.
DOI : 10.1172/JCI119556

B. Marshall, M. Long, J. Piper, T. Yago, and R. Mcever, Direct observation of catch bonds involving cell-adhesion molecules, Nature, vol.77, issue.6936, pp.190-193, 2003.
DOI : 10.1016/0304-4157(86)90016-X

P. Sundd, E. Gutierrez, M. Pospieszalska, H. Zhang, and A. Groisman, Quantitative dynamic footprinting microscopy reveals mechanisms of neutrophil rolling, Nature Methods, vol.103, issue.10, pp.821-824, 2010.
DOI : 10.1080/10739680802462792

R. Mcever and C. Zhu, Rolling Cell Adhesion, Annual Review of Cell and Developmental Biology, vol.26, issue.1, pp.363-396, 2010.
DOI : 10.1146/annurev.cellbio.042308.113238

P. Sundd, E. Gutierrez, E. Koltsova, Y. Kuwano, and S. Fukuda, ???Slings??? enable neutrophil rolling at high shear, Nature, vol.22, issue.7411, pp.399-403, 2012.
DOI : 10.1038/nature11248

URL : http://www.ncbi.nlm.nih.gov/pmc/articles/PMC3433404

J. Takagi, B. Petre, T. Walz, and T. Springer, Global Conformational Rearrangements in Integrin Extracellular Domains in Outside-In and Inside-Out Signaling, Cell, vol.110, issue.5, pp.599-611, 2002.
DOI : 10.1016/S0092-8674(02)00935-2

S. Simon and C. Green, Molecular Mechanics and Dynamics of Leukocyte Recruitment During Inflammation, Annual Review of Biomedical Engineering, vol.7, issue.1, pp.151-185, 2005.
DOI : 10.1146/annurev.bioeng.7.060804.100423

A. Chigaev, T. Buranda, D. Dwyer, E. Prossnitz, and L. Sklar, FRET Detection of Cellular ??4-Integrin Conformational Activation, Biophysical Journal, vol.85, issue.6, pp.3951-3962, 2003.
DOI : 10.1016/S0006-3495(03)74809-7

J. Chan, S. Hyduk, and M. Cybulsky, Detecting rapid and transient upregulation of leukocyte integrin affinity induced by chemokines and chemoattractants, Journal of Immunological Methods, vol.273, issue.1-2, pp.43-52, 2003.
DOI : 10.1016/S0022-1759(02)00417-9

J. Divietro, D. Brown, L. Sklar, R. Larson, and M. Lawrence, Immobilized Stromal Cell-Derived Factor-1?? Triggers Rapid VLA-4 Affinity Increases to Stabilize Lymphocyte Tethers on VCAM-1 and Subsequently Initiate Firm Adhesion, The Journal of Immunology, vol.178, issue.6, pp.3903-3911, 2007.
DOI : 10.4049/jimmunol.178.6.3903

R. Alon, S. Feigelson, E. Manevich, D. Rose, and J. Schmitz, -cytoplasmic domain, The Journal of Cell Biology, vol.95, issue.6, pp.1073-1084, 2005.
DOI : 10.1529/biophysj.104.045690

URL : https://hal.archives-ouvertes.fr/halshs-00480214

J. Rullo, H. Becker, S. Hyduk, J. Wong, and G. Digby, Actin polymerization stabilizes ??4??1 integrin anchors that mediate monocyte adhesion, The Journal of Cell Biology, vol.114, issue.1, pp.115-129, 2012.
DOI : 10.1083/jcb.201107140.dv

L. Boring, J. Gosling, M. Cleary, and I. Charo, Decreased lesion formation in CCR22/2 mice reveals a role for chemokines in the initiation of atherosclerosis, Nature, vol.394, issue.6696, pp.894-897, 1998.
DOI : 10.1038/29788

F. Chow, E. Ozols, D. Nikolic-paterson, R. Atkins, and G. Tesch, Macrophages in mouse type 2 diabetic nephropathy: Correlation with diabetic state and progressive renal injury, Kidney International, vol.65, issue.1, pp.116-128, 2004.
DOI : 10.1111/j.1523-1755.2004.00367.x

R. Alam, J. York, M. Boyars, S. Stafford, and J. Grant, Increased MCP-1, RANTES, and MIP-1alpha in bronchoalveolar lavage fluid of allergic asthmatic patients., American Journal of Respiratory and Critical Care Medicine, vol.153, issue.4, pp.1398-1404, 1996.
DOI : 10.1164/ajrccm.153.4.8616572

C. Abbadie, J. Lindia, A. Cumiskey, L. Peterson, and J. Mudgett, Impaired neuropathic pain responses in mice lacking the chemokine receptor CCR2, Proceedings of the National Academy of Sciences, vol.100, issue.13, pp.7947-7952, 2003.
DOI : 10.1073/pnas.1331358100

J. Kim, S. Gautam, M. Chopp, C. Zaloga, and M. Jones, Expression of monocyte chemoattractant protein-1 and macrophage inflammatory protein-1 after focal cerebral ischemia in the rat, Journal of Neuroimmunology, vol.56, issue.2, pp.127-134, 1995.
DOI : 10.1016/0165-5728(94)00138-E

A. Ferreira, H. Isaacs, J. Hayflick, K. Rogers, and M. Sandig, The p110?? Isoform of PI3K Differentially Regulates ??1 and ??2 Integrin-Mediated Monocyte Adhesion and Spreading and Modulates Diapedesis, Microcirculation, vol.13, issue.6, pp.439-456, 2006.
DOI : 10.1080/10739680600776062

S. Tsuchiya, M. Yamabe, Y. Yamaguchi, Y. Kobayashi, and T. Konno, Establishment and characterization of a human acute monocytic leukemia cell line (THP-1), International Journal of Cancer, vol.42, issue.2, pp.171-176, 1980.
DOI : 10.1002/ijc.2910260208

N. Ashida, H. Arai, M. Yamasaki, and T. Kita, Distinct Signaling Pathways for MCP-1-dependent Integrin Activation and Chemotaxis, Journal of Biological Chemistry, vol.276, issue.19, pp.16555-16560, 2001.
DOI : 10.1074/jbc.M009068200

E. Florin, V. Moy, and H. Gaub, Adhesion forces between individual ligand-receptor pairs, Science, vol.264, issue.5157, pp.415-417, 1994.
DOI : 10.1126/science.8153628

J. Hutter and J. Bechhoefer, Calibration of atomic???force microscope tips, Review of Scientific Instruments, vol.64, issue.7, pp.1868-1873, 1993.
DOI : 10.1063/1.1143970

X. Zhang, E. Wojcikiewicz, and V. Moy, Force Spectroscopy of the Leukocyte Function-Associated Antigen-1/Intercellular Adhesion Molecule-1 Interaction, Biophysical Journal, vol.83, issue.4, pp.2270-2279, 2002.
DOI : 10.1016/S0006-3495(02)73987-8

F. Rico, C. Chu, M. Abdulreda, Y. Qin, and V. Moy, Temperature Modulation of Integrin-Mediated Cell Adhesion, Biophysical Journal, vol.99, issue.5, pp.1387-1396, 2010.
DOI : 10.1016/j.bpj.2010.06.037

D. Tees, R. Waugh, and D. Hammer, A Microcantilever Device to Assess the Effect of Force on the Lifetime of Selectin-Carbohydrate Bonds, Biophysical Journal, vol.80, issue.2, pp.668-682, 2001.
DOI : 10.1016/S0006-3495(01)76047-X

M. Krieg, J. Helenius, C. Heisenberg, and D. Muller, A Bond for a Lifetime: Employing Membrane Nanotubes from Living Cells to Determine Receptor-Ligand Kinetics, Angewandte Chemie International Edition, vol.64, issue.50, pp.9775-9777, 2008.
DOI : 10.1002/anie.200803552

G. Bell, Models for the specific adhesion of cells to cells, Science, vol.200, issue.4342, pp.618-627, 1978.
DOI : 10.1126/science.347575

E. Evans and K. Ritchie, Dynamic strength of molecular adhesion bonds, Biophysical Journal, vol.72, issue.4, pp.1541-1555, 1997.
DOI : 10.1016/S0006-3495(97)78802-7

S. Hyduk, J. Chan, S. Duffy, M. Chen, and M. Peterson, Phospholipase C, calcium, and calmodulin are critical for ??4??1 integrin affinity up-regulation and monocyte arrest triggered by chemoattractants, Blood, vol.109, issue.1, pp.176-184, 2007.
DOI : 10.1182/blood-2006-01-029199

Q. Jing, S. Xin, W. Zhang, P. Wang, and Y. Qin, Lysophosphatidylcholine Activates p38 and p42/44 Mitogen-Activated Protein Kinases in Monocytic THP-1 Cells, but Only p38 Activation Is Involved in Its Stimulated Chemotaxis, Circulation Research, vol.87, issue.1, pp.52-59, 2000.
DOI : 10.1161/01.RES.87.1.52

M. Rief, M. Gautel, F. Oesterhelt, J. Fernandez, and H. Gaub, Reversible Unfolding of Individual Titin Immunoglobulin Domains by AFM, Science, vol.276, issue.5315, pp.1109-1112, 1997.
DOI : 10.1126/science.276.5315.1109

R. Afrin, H. Arakawa, T. Osada, and A. Ikai, Extraction of Membrane Proteins from a Living Cell Surface Using the Atomic Force Microscope and Covalent Crosslinkers, Cell Biochemistry and Biophysics, vol.39, issue.2, pp.101-117, 2003.
DOI : 10.1385/CBB:39:2:101

X. Zhang, S. Craig, H. Kirby, M. Humphries, and V. Moy, Molecular Basis for the Dynamic Strength of the Integrin ??4??1/VCAM-1 Interaction, Biophysical Journal, vol.87, issue.5, pp.3470-3478, 2004.
DOI : 10.1529/biophysj.104.045690

F. Hochmuth, J. Shao, J. Dai, and M. Sheetz, Deformation and flow of membrane into tethers extracted from neuronal growth cones, Biophysical Journal, vol.70, issue.1, pp.358-369, 1996.
DOI : 10.1016/S0006-3495(96)79577-2

M. Sun, J. Graham, B. Hegedus, M. F. Zhang, and Y. , Multiple Membrane Tethers Probed by Atomic Force Microscopy, Biophysical Journal, vol.89, issue.6, pp.4320-4329, 2005.
DOI : 10.1529/biophysj.104.058180

F. Brochard-wyart, N. Borghi, D. Cuvelier, and P. Nassoy, Hydrodynamic narrowing of tubes extruded from cells, Proceedings of the National Academy of Sciences, vol.103, issue.20, pp.7660-7663, 2006.
DOI : 10.1073/pnas.0602012103

R. Afrin and A. Ikai, Force profiles of protein pulling with or without cytoskeletal links studied by AFM, Biochemical and Biophysical Research Communications, vol.348, issue.1, pp.238-244, 2006.
DOI : 10.1016/j.bbrc.2006.07.050

B. Hosu, M. Sun, F. Marga, M. Grandbois, and G. Forgacs, Eukaryotic membrane tethers revisited using magnetic tweezers, Physical Biology, vol.4, issue.2, pp.67-78, 2007.
DOI : 10.1088/1478-3975/4/2/001

M. Sun, N. Northup, F. Marga, T. Huber, and F. Byfield, The effect of cellular cholesterol on membrane-cytoskeleton adhesion, Journal of Cell Science, vol.120, issue.13, pp.2223-2231, 2007.
DOI : 10.1242/jcs.001370

F. Kong, A. Garcia, A. Mould, M. Humphries, and C. Zhu, Demonstration of catch bonds between an integrin and its ligand, The Journal of Cell Biology, vol.42, issue.7, pp.1275-1284, 2009.
DOI : 10.1016/j.molcel.2008.11.018

M. Dembo, D. Torney, K. Saxman, and D. Hammer, The Reaction-Limited Kinetics of Membrane-to-Surface Adhesion and Detachment, Proceedings of the Royal Society B: Biological Sciences, vol.234, issue.1274, pp.55-83, 1988.
DOI : 10.1098/rspb.1988.0038

P. Williams, Analytical descriptions of dynamic force spectroscopy: behaviour of multiple connections, Analytica Chimica Acta, vol.479, issue.1, pp.107-115, 2003.
DOI : 10.1016/S0003-2670(02)01569-6

C. Berlin, R. Bargatze, J. Campbell, U. Von-andrian, and M. Szabo, ??4 integrins mediate lymphocyte attachment and rolling under physiologic flow, Cell, vol.80, issue.3, pp.413-422, 1995.
DOI : 10.1016/0092-8674(95)90491-3

E. Wojcikiewicz, X. Zhang, A. Chen, and V. Moy, Contributions of molecular binding events and cellular compliance to the modulation of leukocyte adhesion, Journal of Cell Science, vol.116, issue.12, pp.2531-2539, 2003.
DOI : 10.1242/jcs.00465

Y. Van-kooyk and C. Figdor, Avidity regulation of integrins: the driving force in leukocyte adhesion, Current Opinion in Cell Biology, vol.12, issue.5, pp.542-547, 2000.
DOI : 10.1016/S0955-0674(00)00129-0

C. Carman and T. Springer, Integrin avidity regulation: are changes in affinity and conformation underemphasized?, Current Opinion in Cell Biology, vol.15, issue.5, pp.547-556, 2003.
DOI : 10.1016/j.ceb.2003.08.003

M. Brown, R. Nijhara, J. Hallam, M. Gignac, and K. Yamada, Chemokine stimulation of human peripheral blood T lymphocytes induces rapid dephosphorylation of ERM proteins, which facilitates loss of microvilli and polarization, Blood, vol.102, issue.12, pp.3890-3899, 2003.
DOI : 10.1182/blood-2002-12-3807

J. Hao, Y. Liu, M. Kruhlak, K. Debell, and B. Rellahan, Phospholipase C???mediated hydrolysis of PIP2 releases ERM proteins from lymphocyte membrane, The Journal of Cell Biology, vol.115, issue.3, pp.451-462, 2009.
DOI : 10.1038/sj.onc.1207768

G. Fu, An Extension of Hertz???s Theory in Contact Mechanics, Journal of Applied Mechanics, vol.74, issue.2, pp.373-374, 2007.
DOI : 10.1115/1.2188017

A. Goldman, R. Cox, and H. Brenner, Slow viscous motion of a sphere parallel to a plane wall???II Couette flow, Chemical Engineering Science, vol.22, issue.4, pp.653-660, 1967.
DOI : 10.1016/0009-2509(67)80048-4

O. Tissot, A. Pierres, C. Foa, M. Delaage, and P. Bongrand, Motion of cells sedimenting on a solid surface in a laminar shear flow, Biophysical Journal, vol.61, issue.1, pp.204-215, 1992.
DOI : 10.1016/S0006-3495(92)81827-1