L. Airoldi and M. Beck, Loss, status and trends for coastal marine habitats of Europe, Marine Biology Boca Raton, vol.45, pp.345-405, 2007.

L. Aragão, Y. Malhi, D. Metcalfe, J. Silva-espejo, E. Jiménez et al., Above- and below-ground net primary productivity across ten Amazonian forests on contrasting soils, Biogeosciences, vol.6, issue.12, pp.2759-2778, 2009.
DOI : 10.5194/bg-6-2759-2009

M. Arain and N. Restrepo-coupe, Net ecosystem production in a temperate pine plantation in southeastern Canada, Agricultural and Forest Meteorology, vol.128, issue.3-4, pp.223-241, 2005.
DOI : 10.1016/j.agrformet.2004.10.003

S. Asare and M. Harlin, SEASONAL FLUCTUATIONS IN TISSUE NITROGEN FOR FIVE SPECIES OF PERENNIAL MACROALGAE IN RHODE ISLAND SOUND1, Journal of Phycology, vol.60, issue.2, pp.254-257, 1983.
DOI : 10.1111/j.0022-3646.1983.00254.x

L. Benedetti-cecchi, PREDICTING DIRECT AND INDIRECT INTERACTIONS DURING SUCCESSION IN A MID-LITTORAL ROCKY SHORE ASSEMBLAGE, Ecological Monographs, vol.70, issue.1, pp.45-72, 2000.
DOI : 10.2307/1940449

M. Bertness, J. Bruno, B. Silliman, and J. Stachowicz, Marine community ecology and conservation, 2014.

M. Bertness, G. Leonard, J. Levine, P. Schmidt, and A. Ingraham, TESTING THE RELATIVE CONTRIBUTION OF POSITIVE AND NEGATIVE INTERACTIONS IN ROCKY INTERTIDAL COMMUNITIES, Ecology, vol.80, issue.8, pp.2711-2726, 1999.
DOI : 10.1086/285461

E. Billard, E. Serrão, G. Pearson, C. Engel, C. Destombe et al., (Fucaceae, Phaeophyceae) and their putative hybrids, European Journal of Phycology, vol.15, issue.4, pp.397-407, 2005.
DOI : 10.1046/j.1365-2540.2001.00968.x
URL : http://sapientia.ualg.pt/bitstream/10400.1/4156/1/Analysis%20of%20sexual%20phenotype%20and%20prezygotic%20fertility%20in%20natural%20populations%20of%20Fucus%20spiralis%2c%20F.%20vesiculosus%20%28Fucaceae%2c%20Phaeophyceae%29%20and%20their%20putative%20hybrids.pdf

T. Binzer and A. Middelboe, From thallus to communities: scale effects and photosynthetic performance in macroalgae communities, Marine Ecology Progress Series, vol.287, pp.65-75, 2005.
DOI : 10.3354/meps287065

T. Binzer and K. Sand-jensen, Production in aquatic macrophyte communities: A theoretical and empirical study of the influence of spatial light distribution, Limnology and Oceanography, vol.47, issue.6, pp.1742-1750, 2002.
DOI : 10.4319/lo.2002.47.6.1742

T. Binzer and K. Sand-jensen, Importance of structure and density of macroalgae communities (Fucus serratus) for photosynthetic production and light utilisation, Marine Ecology Progress Series, vol.235, pp.53-62, 2002.
DOI : 10.3354/meps235053

T. Binzer, K. Sand-jensen, and A. Middelboe, Community photosynthesis of aquatic macrophytes, Limnology and Oceanography, vol.51, issue.6, pp.2722-2733, 2006.
DOI : 10.4319/lo.2006.51.6.2722

G. Chelazzi and M. Vannini, Behavioral adaptation to intertidal life, 1988.
DOI : 10.1007/978-1-4899-3737-7

A. Cheshire, G. Westphalen, A. Wenden, L. Scriven, and R. Bc, Photosynthesis and respiration of phaeophycean-dominated macroalgal communities in summer and winter, Aquatic Botany, vol.55, issue.3, pp.159-170, 1996.
DOI : 10.1016/S0304-3770(96)01071-6

H. Choi and T. Norton, Competition and facilitation between germlings of Ascophyllum nodosum and Fucus vesiculosus, Marine Biology, vol.220, issue.2, pp.525-532, 2005.
DOI : 10.1007/s00227-005-1593-x

H. Christie, K. Norderhaug, and S. Fredriksen, Macrophytes as habitat for fauna, Marine Ecology Progress Series, vol.396, pp.221-233, 2009.
DOI : 10.3354/meps08351

J. Clavier, L. Chauvaud, A. Carlier, E. Amice, M. Van-der-geest et al., Aerial and underwater carbon metabolism of a Zostera noltii seagrass bed in the Banc d???Arguin, Mauritania, Aquatic Botany, vol.95, issue.1, pp.24-30, 2011.
DOI : 10.1016/j.aquabot.2011.03.005
URL : https://hal.archives-ouvertes.fr/hal-00663787

F. Clements, Plant succession: an analysis of the development of vegetation, 1916.
DOI : 10.5962/bhl.title.56234

F. Clements, Plant succession and indicators. The H.W. Wilson company, 1928.

J. Connell and R. Slatyer, Mechanisms of Succession in Natural Communities and Their Role in Community Stability and Organization, The American Naturalist, vol.111, issue.982, pp.1119-1144, 1977.
DOI : 10.1086/283241

R. Cousens, Estimation of Annual Production by the Intertidal Brown Alga Ascophyllum nodosum (L.) Le Jolis, Botanica Marina, vol.27, issue.5, pp.217-227, 1984.
DOI : 10.1515/botm.1984.27.5.217

J. Coyer, G. Hoarau, M. Oudot-le-secq, W. Stam, and J. Olsen, A mtDNA-based phylogeny of the brown algal genus Fucus (Heterokontophyta; Phaeophyta), Molecular Phylogenetics and Evolution, vol.39, issue.1, pp.209-222, 2006.
DOI : 10.1016/j.ympev.2006.01.019

J. Coyer, G. Hoarau, M. Skage, W. Stam, and J. Olsen, (Heterokontophyta; Fucaceae) populations in Iceland and the Faroes: a microsatellite-based assessment, European Journal of Phycology, vol.55, issue.2, pp.235-246, 2006.
DOI : 10.1515/BOT.2002.054

J. Coyer, A. Peters, W. Stam, and J. Olsen, Post-ice age recolonization and differentiation of Fucus serratus L. (Phaeophyceae; Fucaceae) populations in Northern Europe, Molecular Ecology, vol.28, issue.7, pp.1817-1829, 2003.
DOI : 10.1046/j.1365-294X.2002.01510.x

K. Crawley, G. Hyndes, M. Vanderklift, A. Revill, and P. Nichols, Allochthonous brown algae are the primary food source for consumers in a temperate, coastal environment, Marine Ecology Progress Series, vol.376, pp.33-44, 2009.
DOI : 10.3354/meps07810

E. Creis, L. Delage, S. Charton, S. Goulitquer, C. Leblanc et al., Constitutive or Inducible Protective Mechanisms against UV-B Radiation in the Brown Alga Fucus vesiculosus? A Study of Gene Expression and Phlorotannin Content Responses, PLOS ONE, vol.126, issue.6, p.128003, 2015.
DOI : 10.1371/journal.pone.0128003.s001
URL : https://hal.archives-ouvertes.fr/hal-01232402

J. Cucherousset and S. Villéger, Quantifying the multiple facets of isotopic diversity: New metrics for stable isotope ecology, Ecological Indicators, vol.56, pp.152-160, 2015.
DOI : 10.1016/j.ecolind.2015.03.032

G. Darnis, D. Robert, C. Pomerleau, H. Link, P. Archambault et al., Current state and trends in Canadian Arctic marine ecosystems: II. Heterotrophic food web, pelagic-benthic coupling, and biodiversity, Climatic Change, vol.34, issue.1, pp.179-205, 2012.
DOI : 10.1007/s10584-012-0483-8

P. Dauby, A. Khomsi, and J. Bouquegneau, Trophic relationships within intertidal communities of the Brittany coasts: a stable carbon isotope analysis, J Coast Res, vol.14, pp.1202-1212, 1998.

I. Davison, ENVIRONMENTAL EFFECTS ON ALGAL PHOTOSYNTHESIS: TEMPERATURE, Journal of Phycology, vol.26, issue.1, pp.2-8, 1991.
DOI : 10.1111/j.0022-3646.1991.00002.x

I. Davison and G. Pearson, STRESS TOLERANCE IN INTERTIDAL SEAWEEDS, Journal of Phycology, vol.32, issue.2, pp.197-211, 1996.
DOI : 10.1111/j.0022-3646.1996.00197.x

M. Dethier, Disturbance and Recovery in Intertidal Pools: Maintenance of Mosaic Patterns, Ecological Monographs, vol.54, issue.1, pp.99-118, 1984.
DOI : 10.2307/1942457

M. Dethier and S. Williams, Seasonal stresses shift optimal intertidal algal habitats, Marine Biology, vol.57, issue.261, pp.555-567, 2009.
DOI : 10.1007/s00227-008-1107-8

A. Dickson and F. Millero, A comparison of the equilibrium constants for the dissociation of carbonic acid in seawater media, Deep Sea Research Part A. Oceanographic Research Papers, vol.34, issue.10, pp.1733-1743, 1987.
DOI : 10.1016/0198-0149(87)90021-5

J. Dijkstra, J. Boudreau, and M. Dionne, Species-specific mediation of temperature and community interactions by multiple foundation species, Oikos, vol.197, issue.5, pp.646-654, 2012.
DOI : 10.1111/j.1600-0706.2011.19712.x

M. Dring and F. Brown, Photosynthesis of Intertidal Brown Algae During and After Periods of Emersion: A Renewed Search for Physiological Causes of Zonation, Marine Ecology Progress Series, vol.8, pp.301-308, 1982.
DOI : 10.3354/meps008301

C. Duarte and J. Cebrian, The fate of marine autotrophic production, Limnology and Oceanography, vol.41, issue.8, pp.1758-1766, 1996.
DOI : 10.4319/lo.1996.41.8.1758

C. Duarte, J. Middelburg, and N. Caraco, Major role of marine vegetation on the oceanic carbon cycle, Biogeosciences, vol.2, issue.1, pp.1-8, 2005.
DOI : 10.5194/bg-2-1-2005
URL : https://hal.archives-ouvertes.fr/hal-00297509

L. Duarte, F. Rossi, C. Docal, and R. Viejo, Effects of alga Fucus serratus decline on benthic assemblages and trophic linkages at its retreating southern range edge, Marine Ecology Progress Series, vol.527, pp.87-103, 2015.
DOI : 10.3354/meps11248

L. Duarte, R. Viejo, B. Martínez, M. Gómez-gesteira, M. Gallardo et al., Recent and historical range shifts of two canopy-forming seaweeds in North Spain and the link with trends in sea surface temperature, Acta Oecologica, vol.51, pp.1-10, 2013.
DOI : 10.1016/j.actao.2013.05.002

S. Dudgeon and P. Petraitis, SCALE-DEPENDENT RECRUITMENT AND DIVERGENCE OF INTERTIDAL COMMUNITIES, Ecology, vol.82, issue.4, pp.991-1006, 2001.
DOI : 10.2307/1942623
URL : http://scholarworks.csun.edu/bitstream/10211.2/2809/1/DudgeonSteve200104.pdf

J. Ellis, M. Shulman, M. Wood, J. Witman, and S. Lozyniak, REGULATION OF INTERTIDAL FOOD WEBS BY AVIAN PREDATORS ON NEW ENGLAND ROCKY SHORES, Ecology, vol.88, issue.4, pp.853-863, 2007.
DOI : 10.1890/0012-9615(1997)067[0045:EATOPC]2.0.CO;2

T. Farrell, Models and Mechanisms of Succession: An Example From a Rocky Intertidal Community, Ecological Monographs, vol.61, issue.1, pp.95-113, 1991.
DOI : 10.2307/1943001

P. Falkowski, J. Raven, J. Ferreira, and L. Ramos, Aquatic photosynthesis A model for the estimation of annual production rates of macrophyte algae, Aquat Bot, vol.33, pp.53-70, 1989.

J. Fish, S. Fish, M. Flores-molina, D. Thomas, C. Lovazzano et al., A student's guide to the seashore Contreras-Porcia L (2014) Desiccation stress in intertidal seaweeds: effects on morphology, antioxidant responses and photosynthetic performance, Aquat Bot, vol.113, pp.90-99, 1989.

G. Fogg, Algal Adaptation to Stress ? Some General Remarks Algal Adaptation to Environmental Stresses, pp.1-19, 2001.
DOI : 10.1007/978-3-642-59491-5_1

A. Forest, M. Sampei, R. Makabe, H. Sasaki, D. Barber et al., The annual cycle of particulate organic carbon export in Franklin Bay (Canadian Arctic): environmental control and food web implications, J Geophys Res Oceans, vol.113, 2008.

B. Foster, On the Determinants of the Upper Limit of Intertidal Distribution of Barnacles (Crustacea: Cirripedia), The Journal of Animal Ecology, vol.40, issue.1, pp.33-48, 1971.
DOI : 10.2307/3328

B. Fry, Stable Isotope Ecology, 2006.
DOI : 10.1007/0-387-33745-8

J. Gagné, K. Mann, and A. Chapman, Seasonal patterns of growth and storage in Laminaria longicruris in relation to differing patterns of availability of nitrogen in the water, Marine Biology, vol.15, issue.1, pp.91-101, 1982.
DOI : 10.1007/BF00396965

L. Gamfeldt, H. Hillebrand, and P. Jonsson, MULTIPLE FUNCTIONS INCREASE THE IMPORTANCE OF BIODIVERSITY FOR OVERALL ECOSYSTEM FUNCTIONING, Ecology, vol.89, issue.5, pp.1223-1231, 2008.
DOI : 10.1073/pnas.96.4.1463

J. Gattuso, M. Frankignoulle, and R. Wollast, CARBON AND CARBONATE METABOLISM IN COASTAL AQUATIC ECOSYSTEMS, Annual Review of Ecology and Systematics, vol.29, issue.1, pp.405-434, 1998.
DOI : 10.1146/annurev.ecolsys.29.1.405

F. Gévaert, D. Davoult, A. Creach, R. Kling, M. Janquin et al., Carbon and nitrogen content of Laminaria saccharina in the eastern English Channel: biometrics and seasonal variations, Journal of the Marine Biological Association of the UK, vol.81, issue.05, pp.727-734, 2001.
DOI : 10.1017/S0025315401004532

P. D. Giorgio and P. Williams, Respiration in aquatic ecosystems, 2005.

C. Golléty, Fonctionnement (métabolisme et réseau trophique) d'un système intertidal rocheux abrité, la zone à Ascophyllum nodosum, relation avec la biodiversité algale et animale, 2008.

C. Golléty and T. Crowe, Contribution of biofilm to ecosystem functioning in rock pools with different macroalgal assemblages, Marine Ecology Progress Series, vol.482, pp.69-79, 2013.
DOI : 10.3354/meps10238

C. Golléty, P. Riera, and D. Davoult, Complexity of the food web structure of the Ascophyllum nodosum zone evidenced by a ??13C and ??15N study, Journal of Sea Research, vol.64, issue.3, pp.304-312, 2010.
DOI : 10.1016/j.seares.2010.04.003

M. Gómez, F. Barreiro, J. López, M. Lastra, and R. De-la-huz, Deposition patterns of algal wrack species on estuarine beaches, Aquatic Botany, vol.105, pp.25-33, 2013.
DOI : 10.1016/j.aquabot.2012.12.001

I. Gomez and C. Wiencke, Seasonal changes in C, N and major organic compounds and their significance to morpho-functional processes in the endemic Antarctic brown alga Ascoseira mirabilis, Polar Biology, vol.19, issue.2, pp.115-124, 1998.
DOI : 10.1007/s003000050222

M. Goulden, A. Mcmillan, G. Winston, A. Rocha, K. Manies et al., Patterns of NPP, GPP, respiration, and NEP during boreal forest succession, Global Change Biology, vol.15, issue.2, pp.855-871, 2011.
DOI : 10.1111/j.1365-2486.2010.02274.x

D. Häder and F. Figueroa, Photoecophysiology of Marine Macroalgae, Photochemistry and Photobiology, vol.46, issue.1, pp.1-14, 1997.
DOI : 10.1007/BF01313484

D. Hanelt, K. Huppertz, and W. Nultsch, Daily course of photosynthesis and photoinhibition in marine macroalgae investigated in the laboratory and field, Marine Ecology Progress Series, vol.97, pp.31-37, 1993.
DOI : 10.3354/meps097031

R. Haring, M. Dethier, and S. Williams, Desiccation facilitates wave-induced mortality of the intertidal alga Fucus gardneri, Marine Ecology Progress Series, vol.232, pp.75-82, 2002.
DOI : 10.3354/meps232075

R. Hartnoll and S. Hawkins, Patchiness and fluctuations on moderately exposed rocky shores, Ophelia, vol.35, issue.1, pp.53-63, 1985.
DOI : 10.1080/00785236.1985.10426619

B. Hatcher, A. Chapman, and K. Mann, An annual carbon budget for the kelp Laminaria longicruris, Marine Biology, vol.156, issue.1, pp.85-96, 1977.
DOI : 10.1007/BF00386909

S. Hawkins, The influence of season and barnacles on the algal colonization of Patella vulgata exclusion areas, Journal of the Marine Biological Association of the United Kingdom, vol.56, issue.01, pp.1-15, 1981.
DOI : 10.2307/1948498

S. Hawkins and R. Hartnoll, Factors determining the upper limits of intertidal canopy-forming algae, Marine Ecology Progress Series, vol.20, pp.265-271, 1985.
DOI : 10.3354/meps020265

S. Hawkins, D. Watson, A. Hill, S. Harding, M. Kyriakides et al., A COMPARISON OF FEEDING MECHANISMS IN MICROPHAGOUS, HERBIVOROUS, INTERTIDAL, PROSOBRANCHS IN RELATION TO RESOURCE PARTITIONING, Journal of Molluscan Studies, vol.55, issue.2, pp.151-165, 1989.
DOI : 10.1093/mollus/55.2.151

C. Hubas, D. Davoult, T. Cariou, and L. Artigas, Factors controlling benthic metabolism during low tide along a granulometric gradient in an intertidal bay, 2006.

K. Huppertz, D. Hanelt, and W. Nultsch, Photoinhibition of photosynthesis in the marine brown algae Fucus serratus as studied in field experiments, Marine Ecology Progress Series, vol.66, pp.175-182, 1990.
DOI : 10.3354/meps066175

P. Kersen, J. Kotta, M. Bu?as, N. Kolesova, and Z. De?ere, Epiphytes and associated fauna on the brown alga Fucus vesiculosus in the Baltic and the North Seas in relation to different abiotic and biotic variables, Marine Ecology, vol.417, issue.Suppl. 1, pp.87-95, 2011.
DOI : 10.1111/j.1439-0485.2010.00418.x

J. Kim and R. Dewreede, Effects of size and season of disturbance on algal patch recovery in a rocky intertidal community, Marine Ecology Progress Series, vol.133, pp.217-228, 1996.
DOI : 10.3354/meps133217

R. King and W. Schramm, Determination of photosynthetic rates for the marine algae Fucus vesiculosus and Laminaria digitata, Marine Biology, vol.25, issue.3, pp.209-213, 1976.
DOI : 10.1007/BF00387605

J. Kirk, Light and photosynthesis in aquatic ecosystems, 1994.
DOI : 10.1017/cbo9781139168212

M. Knight and P. M. , A Biological Study of Fucus Vesiculosus L. and F. Serratus L., Journal of the Marine Biological Association of the United Kingdom, vol.xxvii, issue.02, pp.439-514, 1950.
DOI : 10.1017/S0025315400055454

K. Krumhansl and R. Scheibling, Production and fate of kelp detritus, Marine Ecology Progress Series, vol.467, pp.281-302, 2012.
DOI : 10.3354/meps09940

M. Lamote and L. Johnson, Temporal and spatial variation in the early recruitment of fucoid algae: the role of microhabitats and temporal scales, Marine Ecology Progress Series, vol.368, pp.93-102, 2008.
DOI : 10.3354/meps07592

M. Lamote, L. Johnson, and Y. Lemoine, Photosynthetic responses of an intertidal alga to emersion: The interplay of intertidal height and meteorological conditions, Journal of Experimental Marine Biology and Ecology, vol.428, pp.16-23, 2012.
DOI : 10.1016/j.jembe.2012.05.021
URL : https://hal.archives-ouvertes.fr/hal-00823519

L. Hir, M. Hily, and C. , Macrofaunal diversity and habitat structure in intertidal boulder fields, Biodiversity & Conservation, vol.55, issue.1, pp.233-250, 2005.
DOI : 10.1007/s10531-005-5046-0

J. Leclerc, P. Riera, C. Leroux, L. Lévêque, and D. Davoult, Temporal variation in organic matter supply in kelp forests: linking structure to trophic functioning, Marine Ecology Progress Series, vol.494, pp.87-105, 2013.
DOI : 10.3354/meps10564
URL : https://hal.archives-ouvertes.fr/hal-01255950

W. Lee, X. Zhang, C. Van-baalen, and C. Arnold, Feeding and reproductive performance of the harpacticoid Tisbe carolinensis (Copepoda, Crustacea) in four algal cultures, Marine Ecology Progress Series, vol.24, pp.273-279, 1985.
DOI : 10.3354/meps024273

P. Legendre, lmodel2: Model II Regression. R package version 1, pp.7-9, 2014.

M. Lehmann, S. Bernasconi, A. Barbieri, and J. Mckenzie, Preservation of organic matter and alteration of its carbon and nitrogen isotope composition during simulated and in situ early sedimentary diagenesis, Geochimica et Cosmochimica Acta, vol.66, issue.20, pp.3573-3584, 2002.
DOI : 10.1016/S0016-7037(02)00968-7

A. Lehvo, S. Bäck, and M. Kiirikki, Growth of Fucus vesiculosus L. (Phaeophyta) in the Northern Baltic Proper: Energy and Nitrogen Storage in Seasonal Environment, Botanica Marina, vol.44, issue.4, pp.345-350, 2001.
DOI : 10.1515/BOT.2001.044

E. Lewis, D. Wallace, and L. Allison, Program developed for CO 2 system calculations. Carbon Dioxide Information Analysis Center, managed by Lockheed Martin Energy Research Corporation for the US Department of Energy Tennessee Littler MM, Littler DS (1980) The evolution of thallus form and survival strategies in benthic marine macroalgae: field and laboratory tests of a functional form model, Am Nat, vol.116, pp.25-44, 1998.

Á. López-urrutia, S. Martin, E. Harris, R. Irigoien, and X. , Scaling the metabolic balance of the oceans, Proceedings of the National Academy of Sciences, vol.23, issue.12, pp.8739-8744, 2006.
DOI : 10.1093/plankt/23.12.1361

D. Loucks, E. Beek, . Van, and J. Stedinger, Water resources systems planning and management: an introduction to methods, models and applications, 2005.
DOI : 10.1007/978-3-319-44234-1

J. Lubchenco, Plant Species Diversity in a Marine Intertidal Community: Importance of Herbivore Food Preference and Algal Competitive Abilities, The American Naturalist, vol.112, issue.983, pp.23-39, 1978.
DOI : 10.1086/283250

J. Lubchenco, Algal Zonation in the New England Rocky Intertidal Community: An Experimental Analysis, Ecology, vol.61, issue.2, pp.333-344, 1980.
DOI : 10.2307/1935192

J. Lubchenco, Littornia and Fucus: Effects of Herbivores, Substratum Heterogeneity, and Plant Escapes During Succession, Ecology, vol.64, issue.5, pp.1116-1123, 1983.
DOI : 10.2307/1937822

K. Lüning, Seaweeds: their environment, biogeography, and ecophysiology, 1990.

S. Maberly and T. Madsen, Contribution of air and water to the carbon balance of Fucus spiralis, Marine Ecology Progress Series, vol.62, pp.175-183, 1990.
DOI : 10.3354/meps062175

T. Madsen and S. Maberly, A COMPARISON OF AIR AND WATER AS ENVIRONMENTS FOR PHOTOSYNTHESIS BY THE INTERTIDAL ALGA FUCUS SPIRALIS (PHAEOPHYTA)1, Journal of Phycology, vol.26, issue.1, pp.24-30, 1990.
DOI : 10.1111/j.0022-3646.1990.00024.x

K. Mann, Seaweeds: Their Productivity and Strategy for Growth: The role of large marine algae in coastal productivity is far more important than has been suspected, Science, vol.73, issue.1, pp.975-981, 1973.
DOI : 10.1111/j.1529-8817.1965.tb04579.x

G. Martins, S. Hawkins, R. Thompson, and S. Jenkins, Community structure and functioning in intertidal rock pools: effects of pool size and shore height at different successional stages, Marine Ecology Progress Series, vol.329, pp.43-55, 2007.
DOI : 10.3354/meps329043

M. Mateo, J. Cebrián, K. Dunton, and T. Mutchler, Carbon Flux in Seagrass Ecosystems, Seagrasses: Biology, Ecology and Conservation, pp.159-192, 2006.
DOI : 10.1007/1-4020-2983-7_7

L. Mccook and A. Chapman, Community succession following massive ice-scour on a rocky intertidal shore: recruitment, competition and predation during early, primary succession, Marine Biology, vol.66, issue.4, pp.565-575, 1993.
DOI : 10.1007/BF00349363

K. Mcguinness and A. Underwood, Habitat structure and the nature of communities on intertidal boulders, Journal of Experimental Marine Biology and Ecology, vol.104, issue.1-3, pp.97-123, 1986.
DOI : 10.1016/0022-0981(86)90099-7

B. Mcmeans, K. Mccann, M. Humphries, N. Rooney, and A. Fisk, Food Web Structure in Temporally-Forced Ecosystems, Trends in Ecology & Evolution, vol.30, issue.11, pp.662-672, 2015.
DOI : 10.1016/j.tree.2015.09.001

C. Mehrbach, C. Culberson, J. Hawley, and R. Pytkowicz, MEASUREMENT OF THE APPARENT DISSOCIATION CONSTANTS OF CARBONIC ACID IN SEAWATER AT ATMOSPHERIC PRESSURE1, Limnology and Oceanography, vol.18, issue.6, pp.897-907, 1973.
DOI : 10.4319/lo.1973.18.6.0897

L. Mesnildrey, C. Jacob, K. Frangoudes, M. Reunavot, and M. Lesueur, La filière des macroalgues en France, 2012.

A. Middelboe and T. Binzer, Importance of canopy structure on photosynthesis in single- and multi-species assemblages of marine macroalgae, Oikos, vol.27, issue.2, pp.422-432, 2004.
DOI : 10.1111/j.0030-1299.2004.13345.x

A. Middelboe, K. Sand-jensen, and T. Binzer, Highly predictable photosynthetic production in natural macroalgal communities from incoming and absorbed light, Oecologia, vol.327, issue.3, pp.464-476, 2006.
DOI : 10.1007/s00442-006-0526-9

A. Migné, D. Davoult, N. Spilmont, D. Menu, G. Boucher et al., A closed-chamber CO 2 -flux method for estimating intertidal primary production and respiration under emersed conditions, Mar Biol, vol.140, pp.865-869, 2002.

A. Migné, G. Delebecq, D. Davoult, N. Spilmont, D. Menu et al., Photosynthetic activity and productivity of intertidal macroalgae: In situ measurements, from thallus to community scale, Aquatic Botany, vol.123, pp.6-12, 2015.
DOI : 10.1016/j.aquabot.2015.01.005

A. Migné, C. Golléty, and D. Davoult, Effect of canopy removal on a rocky shore community metabolism and structure, Marine Biology, vol.94, issue.2, pp.449-457, 2015.
DOI : 10.1007/s00227-014-2592-6

A. Migné, N. Spilmont, and D. Davoult, In situ measurements of benthic primary production during emersion: seasonal variations and annual production in the Bay of Somme (eastern English Channel, France), Continental Shelf Research, vol.24, issue.13-14, pp.1437-1449, 2004.
DOI : 10.1016/j.csr.2004.06.002

R. Miller and H. Page, Kelp as a trophic resource for marine suspension feeders: a review of isotope-based evidence, Marine Biology, vol.17, issue.7, pp.1391-1402, 2012.
DOI : 10.1007/s00227-012-1929-2

R. Miller, D. Reed, and M. Brzezinski, Community structure and productivity of subtidal turf and foliose algal assemblages, Marine Ecology Progress Series, vol.388, pp.1-11, 2009.
DOI : 10.3354/meps08131

K. Moebus, K. Johnson, and J. Sieburth, Rehydration of desiccated intertidal brown algae: Release of dissolved organic carbon and water uptake, Marine Biology, vol.32, issue.2, pp.127-134, 1974.
DOI : 10.1007/BF00388882

M. Molina-montenegro, A. Muoz, E. Badano, B. Morales, K. Fuentes et al., Positive associations between macroalgal species in a rocky intertidal zone and their effects on the physiological performance of Ulva lactuca, Marine Ecology Progress Series, vol.292, pp.173-180, 2005.
DOI : 10.3354/meps292173

M. Molis, J. Koerner, Y. Ko, J. Kim, and M. Wahl, Inducible responses in the brown seaweed Ecklonia cava: the role of grazer identity and season, Journal of Ecology, vol.283, issue.1, pp.243-249, 2006.
DOI : 10.1078/1439-1791-00135

J. Neiva, E. Serrão, J. Assis, G. Pearson, J. Coyer et al., Climate Oscillations, Range Shifts and Phylogeographic Patterns of North Atlantic Fucaceae, pp.279-308, 2016.
DOI : 10.1007/978-94-017-7534-2_11
URL : https://hal.archives-ouvertes.fr/hal-01262364

K. Nicastro, G. Zardi, S. Teixeira, J. Neiva, E. Serrão et al., Shift happens: trailing edge contraction associated with recent warming trends threatens a distinct genetic lineage in the marine macroalga Fucus vesiculosus, BMC Biology, vol.11, issue.1, 2013.
DOI : 10.1093/bioinformatics/btm233

L. Noël, J. Griffin, P. Moschella, S. Jenkins, R. Thompson et al., Changes in Diversity and Ecosystem Functioning During Succession, pp.213-223, 2009.
DOI : 10.1007/b76710_15

M. Nordström, K. Aarnio, and E. Bonsdorff, Temporal variability of a benthic food web: patterns and processes in a low-diversity system, Marine Ecology Progress Series, vol.378, pp.13-26, 2009.
DOI : 10.3354/meps07872

T. Norton, S. Hawkins, N. Manley, G. Williams, and D. Watson, Scraping a living: a review of littorinid grazing, Progress in Littorinid and Muricid Biology, pp.117-138, 1990.

C. Nygård and M. Dring, from the Baltic and Irish Seas, European Journal of Phycology, vol.30, issue.3, pp.253-262, 2008.
DOI : 10.1111/j.0022-3646.1983.00092.x

O. 'brien, A. Morris, L. Keough, and M. , Rapid invertebrate responses to macroalgal wrack: two novel field experiments on intertidal mudflats in Southern Australia, Mar Ecol In press, 2016.

R. Oswald, N. Telford, R. Seed, and C. Happeywood, The effect of encrusting bryozoans on the photosynthetic activity of Fucus serratus L., Estuarine, Coastal and Shelf Science, vol.19, issue.6, pp.697-702, 1984.
DOI : 10.1016/0272-7714(84)90024-6

V. Ouisse, Production primaire et respiration des communautés d'herbiers à zostères : rôle dans le cycle du carbone en milieu côtier, 2010.

V. Ouisse, A. Migné, and D. Davoult, Seasonal variations of community production, respiration and biomass of different primary producers in an intertidal Zostera noltii bed (Western English Channel, France), Hydrobiologia, vol.27, issue.1, pp.3-11, 2010.
DOI : 10.1007/s10750-010-0254-3
URL : https://hal.archives-ouvertes.fr/hal-01255979

V. Ouisse, A. Migné, and D. Davoult, Community-level carbon flux variability over a tidal cycle in Zostera marina and Z. noltii beds, Marine Ecology Progress Series, vol.437, pp.79-87, 2011.
DOI : 10.3354/meps09274
URL : https://hal.archives-ouvertes.fr/hal-01255964

V. Ouisse, P. Riera, A. Migné, C. Leroux, and D. Davoult, Food web analysis in intertidal Zostera marina and Zostera noltii communities in winter and summer, Marine Biology, vol.136, issue.1, pp.165-175, 2011.
DOI : 10.1007/s00227-011-1796-2
URL : https://hal.archives-ouvertes.fr/hal-01255957

C. Oviatt, A. Keller, and L. Reed, Annual Primary Production in Narragansett Bay with no Bay-Wide Winter???Spring Phytoplankton Bloom, Estuarine, Coastal and Shelf Science, vol.54, issue.6, pp.1013-1026, 2002.
DOI : 10.1006/ecss.2001.0872

R. Paine, Trophic Control of Production in a Rocky Intertidal Community, Science, vol.296, issue.5568, pp.736-739, 2002.
DOI : 10.1126/science.1069811

A. Parnell, R. Inger, S. Bearhop, and A. Jackson, Source Partitioning Using Stable Isotopes: Coping with Too Much Variation, PLoS ONE, vol.5, issue.3, p.9672, 2010.
DOI : 10.1371/journal.pone.0009672.g002
URL : http://doi.org/10.1371/journal.pone.0009672

A. Parnell and A. Jackson, Stable Isotope Analysis in R. R package version 4, 2013.

M. Pedersen and J. Borum, Nutrient control of estuarine macroalgae:growth strategy and the balance between nitrogen requirements and uptake, Marine Ecology Progress Series, vol.161, pp.155-163, 1997.
DOI : 10.3354/meps161155

O. Pedersen, T. Colmer, and K. Sand-jensen, Underwater Photosynthesis of Submerged Plants ??? Recent Advances and Methods, Frontiers in Plant Science, vol.4, 2013.
DOI : 10.3389/fpls.2013.00140
URL : http://doi.org/10.3389/fpls.2013.00140

B. Peterson and B. Fry, Stable Isotopes in Ecosystem Studies, Annual Review of Ecology and Systematics, vol.18, issue.1, pp.293-320, 1987.
DOI : 10.1146/annurev.es.18.110187.001453

J. Phillips and C. Hurd, Nitrogen ecophysiology of intertidal seaweeds from New Zealand: N uptake, storage and utilisation in relation to shore position and season, Marine Ecology Progress Series, vol.264, pp.31-48, 2003.
DOI : 10.3354/meps264031

D. Phillips, R. Inger, S. Bearhop, A. Jackson, J. Moore et al., Best practices for use of stable isotope mixing models in food-web studies, Canadian Journal of Zoology, vol.92, issue.10, pp.823-835, 2014.
DOI : 10.1139/cjz-2014-0127

D. Post, USING STABLE ISOTOPES TO ESTIMATE TROPHIC POSITION: MODELS, METHODS, AND ASSUMPTIONS, Ecology, vol.83, issue.3, pp.703-718, 2002.
DOI : 10.4319/lo.1994.39.5.1030

A. Quadir, P. Harrison, and R. Dewreede, The effects of emergence and submergence on the photosynthesis and respiration of marine macrophytes, Phycologia, vol.18, issue.1, pp.83-88, 1979.
DOI : 10.2216/i0031-8884-18-1-83.1

R. Team, R: A Language and Environment for Statistical Computing. R Foundation for Statistical Computing, 2015.

S. Raberg and L. Kautsky, Grazer identity is crucial for facilitating growth of the perennial brown alga Fucus vesiculosus, Marine Ecology Progress Series, vol.361, pp.111-118, 2008.
DOI : 10.3354/meps07428

D. Raffaelli, S. Hawkins, J. Raven, and C. Hurd, Intertidal ecology Ecophysiology of photosynthesis in macroalgae, Photosynth Res, vol.113, pp.105-125, 1999.

J. Raven, A. Johnston, J. Kübler, R. Korb, S. Mcinroy et al., Mechanistic interpretation of carbon isotope discrimination by marine macroalgae and seagrasses, Functional Plant Biology, vol.29, issue.3, pp.355-378, 2002.
DOI : 10.1071/PP01201

J. Rheuban, P. Berg, and K. Mcglathery, Multiple timescale processes drive ecosystem metabolism in eelgrass (Zostera marina) meadows, Marine Ecology Progress Series, vol.507, pp.1-13, 2014.
DOI : 10.3354/meps10843
URL : http://darchive.mblwhoilibrary.org/bitstream/1912/6866/1/m507p001.pdf

A. Ricciardi and E. Bourget, Global patterns of macroinvertebrate biomass in marine intertidal communities, Marine Ecology Progress Series, vol.185, pp.21-35, 1999.
DOI : 10.3354/meps185021

P. Riera, Les isotopes stables en écologie littorale et côtière : principe, exemples de résultats et perspectives, Océanis, vol.32, pp.163-183, 2006.

P. Riera, C. Escaravage, and C. Leroux, Trophic ecology of the rocky shore community associated with the Ascophyllum nodosum zone (Roscoff, France): A ??13C vs ??15N investigation, Estuarine, Coastal and Shelf Science, vol.81, issue.1, pp.143-148, 2009.
DOI : 10.1016/j.ecss.2008.10.018
URL : https://hal.archives-ouvertes.fr/hal-01255980

P. Riera, L. Stal, and J. Nieuwenhuize, Utilization of food sources by invertebrates in a man-made intertidal ecosystem (Westerschelde, the Netherlands): a 13C and 15N study, Journal of the Marine Biological Association of the UK, vol.84, issue.2, pp.323-326, 2004.
DOI : 10.1017/S002531540400921Xh

C. Rigolet, E. Thiébaut, A. Brind-'amour, and S. Dubois, Investigating isotopic functional indices to reveal changes in the structure and functioning of benthic communities, Functional Ecology, vol.89, issue.10, pp.1350-1360, 2015.
DOI : 10.1111/1365-2435.12444
URL : https://hal.archives-ouvertes.fr/hal-01153349

F. Rindi and M. Guiry, Composition and spatio temporal variability of the epiphytic macroalgal assemblage of Fucus vesiculosus Linnaeus at Clare Island, Mayo, western Ireland, Journal of Experimental Marine Biology and Ecology, vol.311, issue.2, pp.233-252, 2004.
DOI : 10.1016/j.jembe.2004.05.009

A. Rio and J. Cabioch, Apparition du Caulacanthus ustulatus (Rhodophyta, Gigartinales) dans la Manche occidentale, Cryptogam Algol, vol.9, pp.231-234, 1988.

S. Rohde, C. Hiebenthal, M. Wahl, R. Karez, and K. Bischof, (Phaeophyceae) in the Western Baltic: effects of light deficiency and epibionts on growth and photosynthesis, European Journal of Phycology, vol.30, issue.2, pp.143-150, 2008.
DOI : 10.3354/meps058175

N. Rooney, K. Mccann, and J. Moore, A landscape theory for food web architecture, Ecology Letters, vol.289, issue.8, pp.867-881, 2008.
DOI : 10.1111/j.1461-0248.2008.01193.x

C. Sacchi, A. Occhipinti-ambrogi, and D. Voltolina, Recherches sur le spectre trophique compare de Littorina saxatilis (Olivi) et de L. nigrolineata (Gray) (Gastropoda, Prosobranchia) sur la grève de Roscoff. II. Cas de populations vivant au milieu d'algues macroscopiques, Cah Biol Mar, vol.22, pp.83-88, 1981.

K. Sand-jensen, Effect of epiphytes on eelgrass photosynthesis, Aquatic Botany, vol.3, pp.55-63, 1977.
DOI : 10.1016/0304-3770(77)90004-3

B. Santelices, Patterns of reproduction, dispersal and recruitment in seaweeds, Oceanogr Mar Biol Annu Rev, vol.28, pp.177-276, 1990.

G. Sarà, D. Pirro, M. Romano, C. Rumolo, P. Sprovieri et al., Sources of organic matter for intertidal consumers on Ascophyllum-shores (SW Iceland): a multi-stable isotope approach, Helgoland Marine Research, vol.7, issue.4, pp.297-302, 2007.
DOI : 10.1007/s10152-007-0078-3

G. Schaal, P. Riera, and C. Leroux, Microscale variations of food web functioning within a rocky shore invertebrate community, Marine Biology, vol.136, issue.3, pp.623-630, 2011.
DOI : 10.1007/s00227-010-1586-2
URL : https://hal.archives-ouvertes.fr/hal-01255968

G. Schaal, P. Riera, C. Leroux, and J. Grall, A seasonal stable isotope survey of the food web associated to a peri-urban rocky shore, Marine Biology, vol.46, issue.2, pp.283-294, 2010.
DOI : 10.1007/s00227-009-1316-9
URL : https://hal.archives-ouvertes.fr/hal-01255976

T. Schlacher and R. Connolly, Effects of acid treatment on carbon and nitrogen stable isotope ratios in ecological samples: a review and synthesis, Methods in Ecology and Evolution, vol.421, issue.6, pp.541-550, 2014.
DOI : 10.1111/2041-210X.12183

A. Schmidt, M. Coll, T. Romanuk, and H. Lotze, Ecosystem structure and services in eelgrass Zostera marina and rockweed Ascophyllum nodosum habitats, Marine Ecology Progress Series, vol.437, pp.51-68, 2011.
DOI : 10.3354/meps09276

M. Schonbeck and T. Norton, Factors controlling the lower limits of fucoid algae on the shore, Journal of Experimental Marine Biology and Ecology, vol.43, issue.2, pp.131-150, 1980.
DOI : 10.1016/0022-0981(80)90021-0

H. Schubert, S. Sagert, and R. Forster, Evaluation of the different levels of variability in the underwater light field of a shallow estuary, Helgoland Marine Research, vol.55, issue.1, pp.12-22, 2001.
DOI : 10.1007/s101520000064

A. Silva, S. Hawkins, D. Boaventura, E. Brewster, and R. Thompson, Use of the intertidal zone by mobile predators: influence of wave exposure, tidal phase and elevation on abundance and diet, Marine Ecology Progress Series, vol.406, pp.197-210, 2010.
DOI : 10.3354/meps08543

S. Smith, Marine Macrophytes as a Global Carbon Sink, Science, vol.211, issue.4484, pp.838-840, 1981.
DOI : 10.1126/science.211.4484.838

J. Smith, S. Vogt, F. Creedon, B. Lucas, and D. Eernisse, The non-native turf-forming alga Caulacanthus ustulatus displaces space-occupants but increases diversity, Biological Invasions, vol.312, issue.10, pp.2195-2208, 2014.
DOI : 10.1007/s10530-014-0658-5

G. Somero, Thermal Physiology and Vertical Zonation of Intertidal Animals: Optima, Limits, and Costs of Living, Integrative and Comparative Biology, vol.42, issue.4, pp.780-789, 2002.
DOI : 10.1093/icb/42.4.780

W. Sousa, The Role of Disturbance in Natural Communities, Annual Review of Ecology and Systematics, vol.15, issue.1, pp.353-391, 1984.
DOI : 10.1146/annurev.es.15.110184.002033

P. South, S. Lilley, L. Tait, T. Alestra, M. Hickford et al., Transient effects of an invasive kelp on the community structure and primary productivity of an intertidal assemblage, Marine and Freshwater Research, vol.67, issue.1, pp.103-112, 2016.
DOI : 10.1071/MF14211

D. Spiller, J. Piovia-scott, A. Wright, L. Yang, G. Takimoto et al., Marine subsidies have multiple effects on coastal food webs, Ecology, vol.134, issue.5, pp.1424-1434, 2010.
DOI : 10.1890/07-0175.1

J. Stachowicz, Mutualism, Facilitation, and the Structure of Ecological Communities, BioScience, vol.51, issue.3, pp.235-246, 2001.
DOI : 10.1641/0006-3568(2001)051[0235:MFATSO]2.0.CO;2

P. Staehr, J. Testa, W. Kemp, J. Cole, K. Sand-jensen et al., The metabolism of aquatic ecosystems: history, applications, and future challenges, Aquatic Sciences, vol.365, issue.1, pp.15-29, 2012.
DOI : 10.1007/s00027-011-0199-2

D. Stagnol, M. Renaud, and D. Davoult, Effects of commercial harvesting of intertidal macroalgae on ecosystem biodiversity and functioning, Estuarine, Coastal and Shelf Science, vol.130, pp.99-110, 2013.
DOI : 10.1016/j.ecss.2013.02.015
URL : https://hal.archives-ouvertes.fr/hal-01255956

L. Tomanek and H. B. , Physiological Ecology of Rocky Intertidal Organisms: A Synergy of Concepts, Integrative and Comparative Biology, vol.42, issue.4, pp.771-775, 2002.
DOI : 10.1093/icb/42.4.771

A. Underwood, The effects of grazing by gastropods and physical factors on the upper limits of distribution of intertidal macroalgae, Oecologia, vol.60, issue.2, pp.201-213, 1980.
DOI : 10.1007/BF00540127

A. Underwood, Microalgal food and the growth of the intertidal gastropods Nerita atramentosa Reeve and Bembicium nanum (Lamarck) at four heights on a shore, Journal of Experimental Marine Biology and Ecology, vol.79, issue.3, pp.277-291, 1984.
DOI : 10.1016/0022-0981(84)90201-6

A. Underwood, Experimental ecology of rocky intertidal habitats: what are we learning?, Journal of Experimental Marine Biology and Ecology, vol.250, issue.1-2, pp.51-76, 2000.
DOI : 10.1016/S0022-0981(00)00179-9

A. Underwood and P. Jernakoff, The effects of tidal height, wave-exposure, seasonality and rock-pools on grazing and the distribution of intertidal macroalgae in New South Wales, Journal of Experimental Marine Biology and Ecology, vol.75, issue.1, pp.71-96, 1984.
DOI : 10.1016/0022-0981(84)90024-8

R. Vadas, S. Johnson, and T. Norton, Recruitment and mortality of early post-settlement stages of benthic algae, British Phycological Journal, vol.8, issue.3, pp.331-351, 1992.
DOI : 10.2307/1942623

R. Vadas, W. Wright, and B. Beal, Biomass and Productivity of Intertidal Rockweeds (Ascophyllum nodosum LeJolis) in Cobscook Bay, Northeastern Naturalist, vol.11, issue.sp2, pp.123-142, 2004.
DOI : 10.1656/1092-6194(2004)11[123:BAPOIR]2.0.CO;2

K. Van-alstyne, J. Ehlig, and S. Whitman, Feeding preferences for juvenile and adult algae depend on algal stage and herbivore species, Marine Ecology Progress Series, vol.180, pp.179-185, 1999.
DOI : 10.3354/meps180179

M. Vanderklift and D. Bearham, Variation in ??13C and ??15N of kelp is explained by light and productivity, Marine Ecology Progress Series, vol.515, pp.111-121, 2014.
DOI : 10.3354/meps10967

M. Vanderklift and S. Ponsard, Sources of variation in consumer-diet ?15N enrichment: a meta-analysis, Oecologia, vol.136, issue.2, pp.169-182, 2003.
DOI : 10.1007/s00442-003-1270-z

I. Viana, A. Bode, M. Bartholomew, and I. Valiela, Experimental assessment of the macroalgae Ascophyllum nodosum and Fucus vesiculosus for monitoring N sources at different time-scales using stable isotope composition, Journal of Experimental Marine Biology and Ecology, vol.466, pp.24-33, 2015.
DOI : 10.1016/j.jembe.2015.01.014

R. Viejo, F. Arenas, C. Fernández, and M. Gómez, Mechanisms of succession along the emersion gradient in intertidal rocky shore assemblages, Oikos, vol.95, issue.3, pp.376-389, 2008.
DOI : 10.1111/j.2007.0030-1299.16206.x

M. Wahl, Marine Hard Bottom Communities: Patterns, Dynamics, Diversity, and Change, 2009.
DOI : 10.1007/b76710

M. Wahl, V. Jormalainen, B. Eriksson, J. Coyer, M. Molis et al., Stress Ecology in Fucus: Abiotic, Biotic and Genetic Interactions, In: Advances in Marine Biology, pp.37-105, 2011.
DOI : 10.1016/B978-0-12-385536-7.00002-9

M. Wahl, M. Molis, A. Hobday, S. Dudgeon, R. Neumann et al., The responses of brown macroalgae to environmental change from local to global scales: direct versus ecologically mediated effects, Perspectives in Phycology, vol.2, issue.1, pp.11-29, 2015.
DOI : 10.1127/pip/2015/0019

F. Warry, P. Reich, R. Woodland, J. Thomson, R. Nally et al., Nitrogen stable isotope values of large-bodied consumers reflect urbanization of coastal catchments, Marine Ecology Progress Series, vol.542, pp.25-37, 2016.
DOI : 10.3354/meps11543

D. Watson and T. Norton, Dietary preferences of the common periwinkle, Littorinalittorea (L.), Journal of Experimental Marine Biology and Ecology, vol.88, issue.3, pp.193-211, 1985.
DOI : 10.1016/0022-0981(85)90230-8

W. Webb, M. Newton, and D. Starr, Carbon dioxide exchange of Alnus rubra, Oecologia, vol.10, issue.4, pp.281-291, 1974.
DOI : 10.1007/BF00345747

S. Williams and M. Dethier, HIGH AND DRY: VARIATION IN NET PHOTOSYNTHESIS OF THE INTERTIDAL SEAWEED FUCUS GARDNERI, Ecology, vol.86, issue.9, pp.2373-2379, 2005.
DOI : 10.1007/s00227-004-1399-2

K. Wyatt, A. Rober, N. Schmidt, and I. Davison, Effects of desiccation and rewetting on the release and decomposition of dissolved organic carbon from benthic macroalgae, Freshwater Biology, vol.169, issue.2, pp.407-416, 2014.
DOI : 10.1111/fwb.12273

K. Wyatt, R. Stevenson, and M. Turetsky, The importance of nutrient co-limitation in regulating algal community composition, productivity and algal-derived DOC in an oligotrophic marsh in interior Alaska, Freshwater Biology, vol.10, issue.9, pp.1845-1860, 2010.
DOI : 10.1111/j.1365-2427.2010.02419.x

E. Young, M. Dring, G. Savidge, D. Birkett, and J. Berges, Seasonal variations in nitrate reductase activity and internal N pools in intertidal brown algae are correlated with ambient nitrate concentrations, Plant, Cell & Environment, vol.288, issue.6, pp.764-774, 2007.
DOI : 10.1016/0022-0981(84)90015-7

M. Zanden and J. Rasmussen, C trophic fractionation: Implications for aquatic food web studies, Limnology and Oceanography, vol.46, issue.8, pp.2061-2066, 2001.
DOI : 10.4319/lo.2001.46.8.2061