R. W. Briehl, The relation between the oxygen equilibrium and aggregation of subunits in lamprey hemoglobin, J. Biol. Chem, vol.238, pp.2361-2366, 1963.

R. S. Carney, Consideration of the oasis analogy for chemosynthetic communities at Gulf of Mexico hydrocarbon vents. Geo-Mar, Lett, vol.14, pp.149-159, 1994.

S. C. Cary, T. Shank, and J. Stein, Worms bask in extreme temperatues, Nature, vol.391, pp.546-546, 1998.

S. C. Cary, R. D. Vetter, and H. Felbeck, Habitat characterization and nutritional strategies of the endosymbiont-bearing bivalve Lucinoma aequizonata, Marine Ecology Progress Series, vol.55, pp.31-45, 1989.
DOI : 10.3354/meps055031

P. Chevaldonné, Ecologie des cheminées hydrothermales actives, Thèse de Doctorat, p.257, 1996.

P. Chevaldonné, D. Desbruyères, and J. J. Childress, . . . and some even hotter, Nature, vol.359, issue.6396, pp.593-594, 1992.
DOI : 10.1038/359593b0

P. Chevaldonné, D. Desbruyères, and M. L. Haitre, Time-series of temperature from three deep-sea hydrothermal vent sites, Deep Sea Research Part A. Oceanographic Research Papers, vol.38, issue.11, pp.1417-1430, 1991.
DOI : 10.1016/0198-0149(91)90014-7

J. J. Childress and C. R. Fischer, The biology of hydrothermal vent animals: physiology, biochemistry and autotrophic symbioses. Oceanogr, Mar. Biol. Annu. Rev, vol.30, pp.337-441, 1992.

J. J. Childress, R. W. Lee, N. K. Sanders, H. Felbeck, D. R. Oros et al., Inorganic carbon uptake in hydrothermal vent tubeworms facilitated by high environmental pC02, Nature, vol.362, issue.6416, pp.147-149, 1993.
DOI : 10.1038/362147a0

J. J. Childress and T. J. , Metabolic rates of animals from the hydrothermal vents and other deep-sea habitats, Bull. Biol. Soc. Wash, vol.6, pp.249-260, 1985.

G. M. Church and W. Gilbert, Genomic sequencing., Proc. Natl. Acad. Sci. USA, pp.1991-1995, 1984.
DOI : 10.1073/pnas.81.7.1991

J. D. Cline and F. A. Richards, Oxygenation of hydrogen sulfide in seawater at constant salinity, temperature and pH, Environmental Science & Technology, vol.3, issue.9, pp.838-843, 1969.
DOI : 10.1021/es60032a004

J. D. Cooper, M. T. Lewis, and D. C. , Pre-column o-phthalaldehyde derivatization of amino acids and their separation using reversed-phase high-performance liquid chromatography, Journal of Chromatography A, vol.285, pp.490-494, 1984.
DOI : 10.1016/S0021-9673(01)87790-6

J. D. Cooper and D. C. , Fluorescence detection of cystine by o-phthalaldehyde derivatisation and its separation using high-performance liquid chromatography, Journal of Chromatography B: Biomedical Sciences and Applications, vol.227, issue.1, pp.158-161, 1982.
DOI : 10.1016/S0378-4347(00)80365-5

C. J. and R. D. Ballard, Oases of life in the cold abyss, Natl. Geogr, vol.152, pp.441-454, 1977.

E. Dahlhoff, J. O-'brien, G. N. Somero, and R. D. Vetter, Temperature Effects on Mitochondria from Hydrothermal Vent Invertebrates: Evidence for Adaptation to Elevated and Variable Habitat Temperatures, Physiological Zoology, vol.64, issue.6, pp.1490-1508, 1991.
DOI : 10.1086/physzool.64.6.30158226

L. J. Dangott and R. C. Terwilliger, Structural studies of a Branchiopod crustacean (Lepidurus bilobatus) extracellular hemoglobin. Evidence for oxygen binding domains, 1979.

B. J. Davis and A. F. Riggs, Effects of sodium chloride and carbon dioxide on oxygen binding byArtemia hemoglobin, Journal of Comparative Physiology B, vol.193, issue.3, pp.355-361, 1986.
DOI : 10.1007/BF01101098

D. Haas, F. , F. Zal, F. H. Lallier, A. Toulmond et al., Three-dimensional reconstruction of the hexagonal bilayer hemoglobin of the hydrothermal vent tube worm Riftia pachyptila by cryoelectron microscopy, Proteins, vol.26, pp.241-256, 1996.

D. Haas, F. , F. Zal, V. You, F. H. Lallier et al., Three-dimensional Reconstruction by Cryoelectron Microscopy of the Giant Hemoglobin of the Polychaete WormAlvinella pompejana, Journal of Molecular Biology, vol.264, issue.1, pp.111-120, 1996.
DOI : 10.1006/jmbi.1996.0627

P. Dejours, Principles of Comparative Respiratory Physiology, 1981.

D. Desbruyères, P. Crassous, J. F. Grassle, A. Khripounoff, D. Reyss et al., Données écologiques sur un nouveau site d'hydrothermalisme actif de la ride du Pacifique oriental, C. R. Acad. Sci. Paris, sér. III, vol.295, pp.489-494, 1982.

D. Desbruyères, F. Gaill, L. Laubier, and Y. Fouquet, Polychaetous annelids from hydrothermal vent ecosystems: An ecological overview, Bull. Biol. Soc. Wash, vol.6, pp.103-116, 1985.

D. Desbruyères and S. Hourdez, A new species of scale-worm (Polychaeta: Polynoidae), Lepidonotopodium atalantae sp. nov., from the East Pacific Rise at 13°N and 9°50'N. Cah, 2000.

D. Desbruyères and S. Hourdez, A new species of scale-worm (Polychaeta: Polynoidae), Lepidonotopodium jouinae sp. nov., from the Azores Triple Junction on the Mid-Atlantic Ridge, Cah. Biol, 2000.

D. Desbruyères and L. Laubier, Systematics, phylogeny, ecology and distribution of the Alvinellidae (Polycheata) from deep-sea hydrothermal vents, Ophelia, vol.5, pp.31-45, 1991.

S. Desbruyères, Handbook of the deep-sea hydrothermal vent fauna, 1997.

S. Dewilde, M. Blaxter, M. Van-hauwaert, J. Vanfleteren, E. L. Esmans et al., Globin and Globin Gene Structure of the Nerve Myoglobin of Aphrodite aculeata, Journal of Biological Chemistry, vol.271, issue.33, pp.19865-19870, 1996.
DOI : 10.1074/jbc.271.33.19865

H. Eisig, Zur systematik, anatomie und morphologie der Ariciiden nebst Beiträgen zur generellen systematik. Mitt, Zool. Stat. Neapel, vol.21, pp.153-593, 1914.

P. Fauvel, Sur les Ampharétiens (Annélides Polychètes) de la côte occidentale de l'Afrique, Bull. Inst. Océanogr. Monaco, vol.1130, pp.1-8, 1958.

T. Somero and . Soto, Microhabitat variation in the hydrothermal vent mussel Bathymodiolus thermophilus, at Rose Garden vent on the Galápagos rift, Deep-Sea Res, vol.35, pp.1769-1792, 1988.

C. R. Fisher, J. J. Childress, S. A. Macko, and J. M. Brooks, Nutritional interactions in Galapagos Rift hydrothermal vent communities: inferences from stable carbon and nitrogen isotope analyses, Marine Ecology Progress Series, vol.103, pp.45-55, 1994.
DOI : 10.3354/meps103045

S. Carney, E. Joye, and . Mcmullin, Methane ice worms: Hesiocaeca methanicola colonizing fossil fuel reserves, Naturwissenschaften, vol.87, pp.184-187, 2000.

H. Furata, M. Ohe, and A. Kajita, Ligand-dependent polymerization of tetrameric hemoglobin from the blood clam Anadara broughtonii, Biochimica et Biophysica Acta (BBA) - Protein Structure, vol.668, issue.3, pp.448-455, 1981.
DOI : 10.1016/0005-2795(81)90179-3

M. Gabe, Techniques histologiques, 1968.

F. Gaill, The biology of annelid worms from high temperature hydrothermal vent regions, Rev. Aquat. Sci, vol.4, pp.107-137, 1991.

F. Gaill, K. Mann, H. Wiedemann, J. Engel, and R. Timpl, Structural Comparison of Cuticle and Interstitial Collagens from Annelids Living in Shallow Sea-water and at Deep-sea Hydrothermal Vents, Journal of Molecular Biology, vol.246, issue.2, pp.284-294, 1995.
DOI : 10.1006/jmbi.1994.0084

W. A. Gallacher and W. B. Elliott, The formation of pyridine haemochromogen, Biochemical Journal, vol.97, issue.1, pp.187-193, 1965.
DOI : 10.1042/bj0970187

I. Gamenick, B. Vismann, M. K. Grieshaber, and O. Giere, Ecophysiological differentiation of Capitella capitata (Polychaeta). Sibling species from different sulfidic habitats, Marine Ecology Progress Series, vol.175, pp.155-166, 1998.
DOI : 10.3354/meps175155

S. L. Gardiner, Respiratory and feeding appendages. in The Ultrastructure of Polychaeta, pp.37-43, 1988.

S. L. Gardiner, Chapter 2.2. Polychaeta: general organization, integument, musculature, coelom, and vascular system, pp.19-52, 1992.

M. Goodman, J. Pedwaydon, J. Czelusniak, T. Suzuki, T. Gotoh et al., An evolutionary tree for invertebrate globin sequences, Journal of Molecular Evolution, vol.67, issue.3, pp.236-249, 1988.
DOI : 10.1007/BF02100080

B. N. Green, T. Suzuki, T. Gotoh, A. R. Kuchumov, and S. N. Vinogradov, Electrospray Ionization Mass Spectrometric Determination of the Complete Polypeptide Chain Composition of Tylorrhynchus heterochaetus Hemoglobin, Journal of Biological Chemistry, vol.270, issue.31, pp.18209-18211, 1995.
DOI : 10.1074/jbc.270.31.18209

M. K. Grieshaber, I. Hardewig, U. Kreutzer, and H. Pörtner, Physiological and metabolic responses to hypoxia in invertebrates, Rev. Physiol. Biochem. Pharmacol, vol.125, pp.43-147, 1994.
DOI : 10.1007/BFb0030909

W. Y. Kao, P. M. Trewitt, and G. Bergtrom, Intron-containing globin genes in the insect Chironomus thummi, Journal of Molecular Evolution, vol.38, issue.3, pp.241-249, 1994.
DOI : 10.1007/BF00176086

G. Kaule, R. Timpl, F. Gaill, and V. Gunzler, Prolyl hydroxylase activity in tissue homogenates of annelids from deep sea hydrothermal vents, Matrix Biology, vol.17, issue.3, pp.205-212, 1998.
DOI : 10.1016/S0945-053X(98)90059-2

M. C. Kennicutt and R. A. Burke, Stable isotopes: Clues to biological cycling of elements at hydrothermal vents. in Microbiology of Deep -Sea Hydrothermal Vents, Corporate Blvd NW, pp.275-287, 1995.

R. E. Kochevar, J. J. Childress, C. R. Fischer, and E. Minnich, The methane mussel: roles of symbiont and host in the metabolic utilization of methane, Marine Biology, vol.1, issue.2, pp.389-401, 1992.
DOI : 10.1007/BF00356284

R. E. Kochevar, N. S. Govind, and J. J. Childress, Identification and characterization of two carbonic anhydrases from the hydrothermal vent tubeworm Riftia pachyptila Jones. Molec, Mar. Biol. Biotechnol, vol.2, pp.10-19, 1993.

A. Krebs, A. R. Kuchumov, P. K. Sharma, E. H. Braswell, P. Zipper et al., Molecular Shape, Dissociation, and Oxygen Binding of the Dodecamer Subunit of Lumbricus terrestris Hemoglobin, Journal of Biological Chemistry, vol.271, issue.31, pp.18695-18704, 1996.
DOI : 10.1074/jbc.271.31.18695

K. Krogh-rasmussen and R. E. Weber, with special reference to the influences of salinity and temperature, Ophelia, vol.4, issue.2, pp.151-170, 1979.
DOI : 10.1080/00785326.1979.10425496

U. K. Laemmli, Cleavage of Structural Proteins during the Assembly of the Head of Bacteriophage T4, Nature, vol.244, issue.5259, pp.680-685, 1970.
DOI : 10.1038/227680a0

P. Lindroth and K. Mopper, High performance liquid chromatographic determination of subpicomole amounts of amino acids by precolumn fluorescence derivatization with o-phthaldialdehyde, Analytical Chemistry, vol.51, issue.11, pp.1667-1674, 1979.
DOI : 10.1021/ac50047a019

P. F. Lonsdale, Clustering of suspension-feeding macrobenthos near abyssal hydrothermal vents at oceanic spreading centers, Deep Sea Research, vol.24, issue.9, pp.857-863, 1977.
DOI : 10.1016/0146-6291(77)90478-7

O. H. Lowry, N. J. Rosebrough, A. L. Farr, and R. J. Randall, Protein measurement with the Folin phenol reagent, J. Biol. Chem, vol.193, pp.265-275, 1951.

J. Lwebuga-mukasa, The role of elytra in hte movement of water over the surface of Halosydna brevisetosa (Polychaeta: Polynoidae), Bull. So. Calif. Acad. Sci, vol.69, pp.154-160, 1970.

I. R. Macdonald, W. R. Callender, R. A. Burke, J. , S. J. Macdonald et al., Fine-scale distribution of methanotrophic mussels at a Louisiana cold seep, Progress in Oceanography, vol.24, issue.1-4, pp.15-24, 1990.
DOI : 10.1016/0079-6611(90)90016-U

I. R. Macdonald, J. F. Reilly, I. , N. L. Guinasso, J. M. Jr et al., Chemosynthetic Mussels at a Brine-Filled Pockmark in the Northern Gulf of Mexico, Science, vol.248, issue.4959, pp.1096-1099, 1990.
DOI : 10.1126/science.248.4959.1096

. Gabrielle, Gulf of Mexico hydrocarbon seep communities: VI. Patterns in community structure and habitat. Geo-Mar, Lett, vol.10, pp.244-252, 1990.

A. J. Magenheim and J. M. Gieskes, Hydrothermal discharge and alteration in near-surface sediments from the Guaymas Basin, Gulf of California, Geochimica et Cosmochimica Acta, vol.56, issue.6, pp.2329-2338, 1992.
DOI : 10.1016/0016-7037(92)90192-L

C. P. Mangum, Multiple Sites of Gas Exchange, American Zoologist, vol.34, issue.2, pp.184-193, 1994.
DOI : 10.1093/icb/34.2.184

C. P. Mangum, B. R. Woodin, C. Bonaventura, B. Sullivan, and J. Bonaventura, The role of coelomic and vascular hemoglobin in the annelid family terebellidae, Comparative Biochemistry and Physiology Part A: Physiology, vol.51, issue.2, pp.281-294, 1975.
DOI : 10.1016/0300-9629(75)90372-2

A. M. Manning, C. N. Trotman, and W. P. Tate, Evolution of a polymeric globin in the brine shrimp Artemia, Nature, vol.348, issue.6302, pp.653-656, 1990.
DOI : 10.1038/348653a0

D. Martin and T. A. Britayev, Symbiotic Polychaetes: review of the known species, 1998.

P. Martineu, S. K. Juniper, C. R. Fisher, and G. J. Massoth, Sulfide Binding in the Body Fluids of Hydrothermal Vent Alvinellid Polychaetes, Physiological Zoology, vol.70, issue.5, pp.578-588, 1997.
DOI : 10.1086/515864

G. J. Massoth, D. A. Butterfield, J. E. Lupton, R. E. Mc-duff, M. D. Lilley et al., Submarine venting of phase-separated hydrothermal fluids at Axial Volcano, Juan de Fuca Ridge, Nature, vol.340, issue.6236, pp.702-705, 1989.
DOI : 10.1038/340702a0

J. G. Menon and A. J. Arp, Morphological adaptations of the respiratory hindgut of a marine echiuran worm, Journal of Morphology, vol.1, issue.2, pp.131-138, 1992.
DOI : 10.1002/jmor.1052140203

J. G. Menon and A. J. Arp, The Integument of the Marine Echiuran Worm Urechis caupo, The Biological Bulletin, vol.185, issue.3, pp.440-454, 1993.
DOI : 10.2307/1542484

T. Miura, Two New Scale-Worms (Polynoidae, Polychaeta) from the Lau Back-Arc and North Fiji Basins, South Pacific Ocean, Proc. Biol. Soc. Wash, vol.107, pp.532-543, 1994.

T. Miura and D. Desbruyères, Two new species of Opisthotrochopodus (Polychaeta: Polynoidae: Branchinotogluminae) from the Lau and the North Fiji Bac-arc Basins, southwestern Pacific Ocean, Proc Biol Soc Wash, vol.108, pp.583-595, 1995.

T. Miura and J. Hashimoto, Two new branchiate scale-worms (Polynoidae: Polycheata) from the hydrothermal vent of the Okinawa trough and the volcanic seamount off Chichijima island, Proc. Biol. Soc. Wash, vol.104, pp.166-174, 1991.

S. Moore and W. H. Stein, Chromatography of amino acids on sulfonated polystyrene resins, J. Biol. Chem, vol.192, pp.663-681, 1951.

Y. Naito, C. K. Riggs, T. L. Vandergon, and A. F. Riggs, Origin of a "bridge" intron in the gene for a two-domain globin., Proc. Natl. Acad. Sci. USA, pp.6672-6676, 1991.
DOI : 10.1073/pnas.88.15.6672

A. W. Nichol, I. Hendry, D. B. Morell, and P. S. Clezy, Mechanism of formation of sulphhaemoglobin, Biochimica et Biophysica Acta (BBA) - General Subjects, vol.156, issue.1, pp.97-108, 1968.
DOI : 10.1016/0304-4165(68)90108-6

P. Nicholls, The effect of sulphide on cytochromeaa3 Isosteric and allosteric shifts of the reduced ??-peak, Biochimica et Biophysica Acta (BBA) - Bioenergetics, vol.396, issue.1, pp.24-35, 1975.
DOI : 10.1016/0005-2728(75)90186-3

E. R. Nix, C. R. Fisher, J. Vodenichar, and K. M. Scott, Physiological ecology of a mussel with methanotrophic endosymbionts at three hydrocarbon seep sites in the Gulf of Mexico, Marine Biology, vol.73, issue.4, pp.605-617, 1995.
DOI : 10.1007/BF00350682

O. 'brien, J. , and R. D. Vetter, Production of thiosulfate during sulphide oxidation by mitochondria of the symbiont-containing bivalve Solemya reidi, J. Exp. Biol, vol.149, pp.133-148, 1990.

L. C. Oglesby, Salt and water balance. in Physiology of annelids, pp.555-658, 1978.

T. Okazaki, R. Briehl, and J. Wittenberg, The hemoglobin of ascaris perienteric fluid, Biochimica et Biophysica Acta (BBA) - General Subjects, vol.111, issue.2, pp.496-502, 1965.
DOI : 10.1016/0304-4165(65)90059-0

C. M. Park and R. L. Nagel, Sulfhemoglobinemia. Clinical and molecular aspects, N. Engl, 1984.

S. Patel and C. P. Spencer, The Oxidation of Sulphide by the Haem Compounds From the Blood of Arenicola Marina, Journal of the Marine Biological Association of the United Kingdom, vol.2, issue.01, pp.167-175, 1963.
DOI : 10.1016/S0003-2670(00)88494-9

K. G. Paul, H. Theorell, and A. Akeson, The Molar Light Absorption of Pyridine Ferroprotoporphrin (Pyridine Haemochromogen)., Acta Chemica Scandinavica, vol.7, pp.1284-1287, 1953.
DOI : 10.3891/acta.chem.scand.07-1284

B. Pelster and R. E. Weber, The Physiology of the Root Effect, Adv. Comp. Environm. Physiol, vol.8, pp.51-77, 1991.
DOI : 10.1007/978-3-642-75900-0_2

M. Perutz, Mechanisms of cooperativity and allosteric regulation in proteins, Quarterly Reviews of Biophysics, vol.199, issue.02, 1990.
DOI : 10.1038/232197a0

M. H. Pettibone, A new scale worm (Polychaeta: Polynoidae) from the hydrothermal riftarea off Western Mexico at 21°N. Proc, Biol. Soc. Wash, vol.96, pp.392-399, 1983.

M. H. Pettibone, A new scale-worm commensal with deep sea mussels on the Galápagos hydrothermal vent (Polychaeta: Polynoidae), Proc. Biol. Soc. Wash, vol.97, pp.226-239, 1984.

M. H. Pettibone, Two new species of Lepidonotopodium, 1984.

M. H. Pettibone, Additionnal branchiate scale worms (Polychaeta: Polynoidae) from Galápagos hydrothermal vent and rift-area off Western Mexico at 21°N, Proc. Biol. Soc. Wash, vol.98, pp.447-469, 1985.

M. H. Pettibone, An additionnal new scale worm (Polycheata: Polynoidae) from the hydrothermal rift area off western Mexico at 21°N, Proc. Biol. Soc. Wash, vol.98, pp.150-157, 1985.

M. H. Pettibone, New genera and species of deep-sea Macellicephalinae and Harmothoinae (Polycheata: Polynoidae) from the hydrothermal rift areas off Galápagos and Western Mexico at 21°N and from the Santa Catalina Channel, Proc. Biol. Soc. Wash, vol.98, pp.740-757, 1985.

M. H. Pettibone, A new scale-worm commensal with deep-sea mussels in the seep-sites at the Florida escarpment in the Eastern Gulf of Mexico, 1986.

M. H. Pettibone, New species and new records of scaled polycheates (Polycheata: Polynoidae) from hydrothermal vents of the Northeast Pacific Explorer and Juan de Fuca Ridges, Proc. Biol. Soc. Wash, vol.101, pp.192-208, 1988.

M. H. Pettibone, New species of scaled-worms (Polycheata: Polynoidae) from the hydrothermal rift-area of the Mariana back-arc basin in the Western central Pacific, Proc. Biol, 1989.

M. H. Pettibone, Polynoidae and Sigalionidae (Polycheata) from the Guaymas basin, with descriptions of two new species, and additionnal records from hydrothermal vents of the Galapagos rift, 21°N, and seep-sites in the gulf of Mexico (Florida and Louisiana), Proc. Biol, 1989.

M. H. Pettibone, Two new species of Harmothoinae (Polycheata: Polynoidae) from the East Pacific Rise, Proc. Biol. Soc. Wash, vol.102, pp.305-310, 1989.

M. H. Pettibone, New species and new records of scaled polychaetes (Polychaeta : Polynoidae) from the axial seamount caldera of the Juan de Fuca Ridge in the Northeast Pacific and the East Pacific ocean off Northern California, Proc. Biol. Soc. Wash, vol.103, pp.825-838, 1990.

M. H. Pettibone, Polynoid polychaetes associated with a whale skeleton in the bathyal santa catalina basin, Proc. Biol. Soc. Wash, vol.106, pp.678-688, 1993.

M. H. Pettibone, Additional records of polynoid polychaetes from the juan de fuca ridge, Proc. Biol. Soc. Wash, vol.107, pp.609-614, 1994.

H. O. Pörtner, R. G. Boutilier, Y. Tang, and D. P. Toews, Determination of intracellular pH and PCO2 after metabolic inhibiton by fluoride and nitrilotriacetic acid, Respiration Physiology, vol.81, issue.2, pp.255-274, 1990.
DOI : 10.1016/0034-5687(90)90050-9

H. O. Pörtner, L. Peck, S. Zielinsky, and L. Z. Conway, Intracellular pH and energy metabolism in the highly stenothermal Antarctic bivalve Limopsis marionensis as a function of ambient temperature, Polar Biology, vol.22, issue.1, pp.17-30, 1999.
DOI : 10.1007/s003000050386

M. A. Powell and G. N. Somero, ADAPTATIONS TO SULFIDE BY HYDROTHERMAL VENT ANIMALS: SITES AND MECHANISMS OF DETOXIFICATION AND METABOLISM, The Biological Bulletin, vol.171, issue.1, pp.274-290, 1986.
DOI : 10.2307/1541923

M. A. Powell and G. N. Somero, Hydrogen Sulfide Oxidation Is Coupled to Oxidative Phosphorylation in Mitochondria of Solemya reidi, Science, vol.233, issue.4763, pp.563-566, 1986.
DOI : 10.1126/science.233.4763.563

M. S. Roberts, R. C. Terwilliger, and N. B. Terwilliger, Comparison of sea cucumber hemoglobin structures, Comparative Biochemistry and Physiology Part B: Comparative Biochemistry, vol.77, issue.1, pp.237-243, 1984.
DOI : 10.1016/0305-0491(84)90250-5

G. W. Rouse and K. Fauchald, The articulation of annelids, Zoologica Scripta, vol.103, issue.1087, pp.269-301, 1995.
DOI : 10.1006/clad.1993.1001

W. E. Royer, Structures of red blood cell hemoglobins. in Oxygen Carriers in Blood and Tissues, pp.87-116, 1992.

J. Sambrook, E. F. Fritsch, and T. Maniatis, Molecular cloning: a laboratory manual, 1989.

N. K. Sanders and J. J. Childress, The use of single column ion chromatography to measure ion concentrations in invertebrate body fluids, Comparative Biochemistry and Physiology Part A: Physiology, vol.98, issue.1, pp.97-100, 1991.
DOI : 10.1016/0300-9629(91)90583-X

P. M. Sarradin, J. C. Caprais, R. Riso, T. Comtet, and A. Aminot, Brief account of the chemical environment at hydrothermal vent mussel beds on the MAR, Cah Biol Mar, vol.39, pp.253-254, 1998.

P. M. Sarradin, J. C. Caprais, R. Riso, R. Kerouel, and A. Aminot, Chemical environment of the hydrothermal mussel communities in the Lucky Strike and Menez Gwen vent fields, Mid Atlantic ridge, Cah Biol Mar, vol.40, pp.93-104, 1999.

B. C. Schaefer, Revolutions in Rapid Amplification of cDNA Ends: New Strategies for Polymerase Chain Reaction Cloning of Full-Length cDNA Ends, Analytical Biochemistry, vol.227, issue.2, pp.255-273, 1995.
DOI : 10.1006/abio.1995.1279

D. Schiedek, Ecophysiological capability ofMarenzelleria populations inhabiting North Sea estuaries: an overview, Helgol??nder Meeresuntersuchungen, vol.31, issue.3-4, pp.373-382, 1998.
DOI : 10.1007/BF02908911

D. Schiedek, C. Vogan, J. Hardege, and M. Bentley, Marenzelleria cf. wireni (Polychaeta: Spionidae) from the Tay estuary. Metabolic response to severe hypoxia and hydrogen sulphide, Aquatic Ecology, vol.31, issue.2, pp.211-222, 1997.
DOI : 10.1023/A:1009911723907

H. Sick and K. Gersonde, Method for Continuous Registration of O2-Binding Curvesof Hemoproteins by Means of a Diffusion Chamber, Analytical Biochemistry, vol.32, issue.3, pp.362-376, 1969.
DOI : 10.1016/S0003-2697(69)80002-3

R. J. Simpson, M. R. Neuberger, and T. Liu, Complete amino-acid analysis of proteins from a single hydrolysate, J. Biol. Chem, vol.251, pp.1936-1940, 1976.

J. D. Smit, H. Sick, A. Peterhans, and K. Gersonde, Acid Bohr effect of a monomeric haemoglobin from Dicrocoelium dendriticum. Mechanism of the allosteric conformation transition, European Journal of Biochemistry, vol.166, issue.2, pp.231-237, 1986.
DOI : 10.1016/0022-2836(79)90332-2

E. B. Smith, K. M. Scott, E. R. Nix, C. Korte, and C. R. Fisher, Growth and condition of seep mussels (Bathymodiolus childressi) at a Gulf of Mexico Brine Pool, Ecology, vol.81, 2000.

G. N. Somero, A. E. Anderson, and J. J. Childress, Transport, metabolism and detoxification of hydrogen sulfide in animals from sulfide-rich marine environments, CRC Crit. Rev. Aq. Sci, vol.1, pp.591-614, 1989.

V. Storch and G. Alberti, Ultrastructural observations on the gills of polychaetes, Helgol??nder Wissenschaftliche Meeresuntersuchungen, vol.90, issue.1-2, pp.169-179, 1978.
DOI : 10.1007/BF02296995

T. Suzuki, Amino acid sequence of the monomer subunit of the giant multisubunit hemoglobin from the earthworm Pheretima sieboldi, European Journal of Biochemistry, vol.241, issue.1, pp.127-134, 1989.
DOI : 10.1111/j.1432-1033.1989.tb15092.x

T. Suzuki, T. Tagagi, and S. Ohta, N-terminal amino acid sequences of 440 kDa hemoglobins of the deep-sea tube worms, Lamellibrachia sp.1, Lamellibrachia sp.2 and slender vestimentifera gen. sp.1. Evolutionary relationship with annelid hemoglobins, Zool. Sci, vol.10, pp.141-146, 1993.

T. Suzuki, T. Takagi, and S. Ohta, N-terminal amino acid sequence of the deep-sea tube worm haemoglobin remarkably resembles that of annelid haemoglobin, Biochem. J, vol.255, pp.541-545, 1988.

J. P. Truchot and F. H. Lallier, High CO 2 content in hydrothermal vent water at the snake pit area, mid-Atlantic Ridge, Cah Biol Mar, vol.39, pp.153-158, 1998.

V. Tunnicliffe, A. G. Mcarthur, and D. Mchugh, A Biogeographical Perspective of the Deep-Sea Hydrothermal Vent Fauna, Advances In Marine Biology, 1998.
DOI : 10.1016/S0065-2881(08)60213-8

I. Tyuma, Y. Kamigawara, and K. Imai, pH dependence of the shape of the hemoglobin-oxygen equilibrium curve, Biochimica et Biophysica Acta (BBA) - Protein Structure, vol.310, issue.2, pp.317-320, 1973.
DOI : 10.1016/0005-2795(73)90111-6

O. W. Van-assendelft, Spectrophotometry of haemoglobin derivatives (Royal Vangorcum Ltd, 1970.

S. N. Vinogradov, O. H. Kapp, and M. Ohtsuki, The extracellular haemoglobins and chlorocruorins of annelids, J. Harris, 1982.

S. N. Vinogradov, J. M. Shlom, O. H. Kapp, and P. Frossard, The dissociation of annelid extracellular hemoglobins and their quaternary structure, Comparative Biochemistry and Physiology Part B: Comparative Biochemistry, vol.67, issue.1, pp.1-16, 1980.
DOI : 10.1016/0305-0491(80)90262-X

. Trotman, Adventious variability? The amino acid sequences of nonvertebrate globins, 1993.

B. Vismann, Sulfide detoxification and tolerance in Nereis (Hediste) diversicolor and Nereis (Neanthes) virens (Annelida. Polychaeta), Marine Ecology Progress Series, vol.59, pp.229-239, 1990.
DOI : 10.3354/meps059229

S. Völkel and M. K. Grieshaber, Mitochondrial Sulfide Oxidation in Arenicola Marina. Evidence for Alternative Electron Pathways, European Journal of Biochemistry, vol.4, issue.1-2, pp.231-237, 1996.
DOI : 10.1016/0304-4173(73)90002-5

V. Damm and K. L. , Seafloor hydrothermal activity: black smocker chemistry and chimneys, 1990.

R. E. Weber, Respiration. in Physiology of Annelids, pp.369-392, 1978.

R. E. Weber, Respiratory pigments. in Physiology of Annelids, pp.393-437, 1978.

R. E. Weber, Functions of Invertebrate Hemoglobins with Special Reference to Adaptations to Environmental Hypoxia, American Zoologist, vol.20, issue.1, pp.79-101, 1980.
DOI : 10.1093/icb/20.1.79

R. E. Weber, Cationic control of O2 affinity in lugworm erythrocruorin, Nature, vol.32, issue.5821, pp.386-387, 1981.
DOI : 10.1038/292386a0

R. E. Weber and J. Baldwin, Blood and erythrocruorin of the giant earthworm, Megascolides australis: respiratory characterisitcs and evidence for CO 2 facilitation of O 2 binding, Mol. Physiol, vol.7, pp.93-106, 1985.

R. E. Weber, G. Braunitzer, T. Kleinschmidt, C. Iiß, C. Iii et al., Functional multiplicity and structural correlations in the hemoglobin system of larvae of Chironomus thummi thummi (Insecta, Diptera): Hb components CTT I, Biochem. Physiol, vol.80, pp.747-753, 1985.

R. E. Weber, C. P. Mangum, H. Steinman, C. Bonaventura, B. Sullivan et al., Hemoglobins of two terebellid polychaetes: Enoplobranchus sanguineus and Amphitrite ornata, Comparative Biochemistry and Physiology Part A: Physiology, vol.56, issue.2, pp.179-187, 1977.
DOI : 10.1016/0300-9629(77)90182-7

R. M. Wells and R. P. Dales, Haemoglobin function in Terebella lapidaria L., an intertidal terebellid polychaete, Journal of the Marine Biological Association of the United Kingdom, vol.46, issue.01, pp.211-220, 1975.
DOI : 10.1016/0305-0491(73)90194-6

R. M. Wells, M. J. Hudson, and T. Brittain, Function of the hemoglobin and the gas bubble in the backswimmer Anisops assimilis (Hemiptera: Notonectidae), J. Comp. Biochem, 1981.