New faunistical and taxonomic data on oribatid mites (Acari: Oribatida) of Ethiopia, with description of two new species of the superfamily Oripodoidea

The present study is based on oribatid mite material collected in Ethiopia in 2019. A list of identified taxa, including 11 species from 10 genera and nine families, is provided; of these, six species, four genera and two families are recorded for the first time in the fauna of this country. Two new species of the superfamily Oripodoidea are described: Pilobates wachteli n. sp. (Haplozetidae); and Zetorchella robertbecki n. sp. (Caloppiidae).


Introduction
This work is based on oribatid mite material (Acari, Oribatida) which was collected during a field trip in southwestern Ethiopia in 2019. The primary goal of our paper is to present a list of the identified taxa with notes on new findings (new records).
The secondary goal of our paper is to describe and illustrate two new species, one belonging to the genus Pilobates Balogh, 1960 (family Haplozetidae), the other to Zetorchella Berlese, 1916 (family Caloppiidae).
Pilobates was proposed by Balogh (1960) with Protoribates pilosellus Balogh, 1958 as type species. The genus comprises seven species which are distributed in the Afrotropical region (mostly Madagascar) and in the Mediterranean (Subías 2021). The revised generic diagnosis was presented by Ermilov (2020). An identification key to all species of Pilobates was provided earlier by Ermilov and Starý (2020). Zetorchella was proposed by Berlese (1916a) with Chaunoproctus cancellatus Pearce, 1906 as type species. The genus comprises 24 species which are distributed in the Afrotropical and Oriental regions, and Guyana (Subías 2021). The revised generic diagnosis and an identification key to species of Zetorchella of the Afrotropical region were presented by Ermilov and N'Dri (2019).

Granuloppiidae
Hammerella sp.: 1 ex. This presumably is a new species; however, we do not describe it here due to the presence of only one specimen. New record of the family and genus in Ethiopia.
Bipassalozetes sp.: 1 ex. This presumably is a new species; however, we do not describe it here due to the presence of only one specimen.
Remarks. The Ethiopian specimens completely correspond morphologically to specimens from South Africa (Ermilov et al. 2019) except for the slightly wavy end of the interlamellar seta (versus straight). We believe this is intraspecific variability.

Haplozetidae
Pilobates wachteli n. sp.: 2 ex. New record of the genus in Ethiopia.
Hence, we found 11 species from 10 genera and nine families. Of these: four species are new to science; six species, four genera and two families are recorded in the fauna of Ethiopia for the first time.
Legs (Figs 3e-h) -Tridactylous. Median claw distinctly thicker than lateral claws, all slightly barbed on dorsal side. Tibia I with small dorsodistal apophysis bearing solenidia φ 1 and φ 2 . Tibiae I and II with triangular tubercle proximoventrally. All femora rounded distoventrally. Trochanter IV rounded distodorsally (without tooth). Dorsoparaxial porose area on femora I-IV and on trochanters III, IV well visible; proximoventral porose area on tarsi I-IV and   Table 1. Famulus on tarsus I short, erect, slightly swollen distally, inserted between solenidion ω 2 and seta ft". Seta s on tarsus I eupathidial, located before setae a' and a". Seta l" on genu I located on small apophysis. Solenidion ω 1 on tarsi I, ω 1 and ω 2 on tarsus II and σ on genu III bacilliform, other solenidia setiform.
Type deposition -The holotype is deposited in the collection of the ZSM; one paratype is deposited in the collection of the TSUMZ.
Etymology -The specific name is dedicated to our friend and colleague, the coleopterologist and all-around biologist Franz Wachtel (Grünwald, Germany) for his expertise and assistance by collecting the Ethiopian mites.
Type deposition -The holotype is deposited in the collection of the ZSM; three paratypes are deposited in the collection of the SMNH; 24 paratypes are deposited in the collection of the TSUMZ.
Etymology -The specific name is dedicated to our friend and colleague, the coleopterologist Robert Beck (Munich, Germany) for his more than generous provision of all necessary facilities during this field trip.