A. P. Anzidei, G. Cavarretta, P. Gioia, M. Mussi, and . Palombo, Tools from elephant bones at la Polledrara di Cecanibbio and Rebibbia-Casal de'Pazzi, The World of Elephants. Proceedings of the First International Congress. Consiglio Nazionale Delle Ricerche, pp.415-418, 2001.

A. Arroyo and I. De-la-torre, Pounding tools in HWK EE and EF-HR (Olduvai Gorge, Tanzania): Percussive activities in the Oldowan-Acheulean transition, Journal of Human Evolution, vol.120, pp.402-421, 2018.

L. Backwell and F. D'errico, Early hominid bone tools from Drimolen, South Africa, Journal of Archaeological Science, vol.35, issue.11, pp.2880-2894, 2008.
URL : https://hal.archives-ouvertes.fr/halshs-00445523

L. R. Backwell and F. D'errico, Evidence of termite foraging by Swartkrans early hominids, Proceedings of the National Academy of Sciences, vol.98, issue.4, pp.1358-1363, 2001.

L. R. Backwell and F. Errico, The first use of bone tools: a reappraisal of the evidence from Olduvai Gorge, Tanzania. Palaeontol. Afr, vol.40, pp.95-158, 2004.
URL : https://hal.archives-ouvertes.fr/halshs-00452494

A. K. Behrensmeyer, Taphonomic and ecologic information from bone weathering, Paleobiology, vol.4, issue.2, pp.150-162, 1978.

F. Bibi, M. Pante, A. Souron, K. Stewart, S. Varela et al., Paleoecology of the Serengeti during the Oldowan-Acheulean transition at Olduvai Gorge, Tanzania: The mammal and fish evidence, Journal of Human Evolution, vol.120, pp.48-75, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02871850

I. Biddittu, P. Celletti, G. Cavarretta, P. Gioia, M. Mussi et al., Plio-Pleistocene Proboscidea and Lower Palaeolithic bone industry of Southern Latium (Italy), The World of Elephants. Proceedings of the First International Congress. Consiglio Nazionale Delle Ricerche, pp.91-96, 2001.

R. Blasco, J. Rosell, F. Cuartero, J. Fernández-peris, A. Gopher et al., Using Bones to Shape Stones: MIS 9 Bone Retouchers at Both Edges of the Mediterranean Sea, PLoS ONE, vol.8, issue.10, p.e76780, 2013.

R. J. Blumenschine, C. W. Marean, and S. D. Capaldo, Blind Tests of Inter-analyst Correspondence and Accuracy in the Identification of Cut Marks, Percussion Marks, and Carnivore Tooth Marks on Bone Surfaces, Journal of Archaeological Science, vol.23, issue.4, pp.493-507, 1996.

R. J. Blumenschine, K. A. Prassack, C. D. Kreger, and M. C. Pante, Carnivore tooth-marks, microbial bioerosion, and the invalidation of Domínguez-Rodrigo and Barba's (2006) test of Oldowan hominin scavenging behavior, Journal of Human Evolution, vol.53, issue.4, pp.420-426, 2007.

A. Bouzouggar, L. T. Humphrey, N. Barton, S. A. Parfitt, L. Clark-balzan et al., 90,000 year-old specialised bone technology in the Aterian Middle Stone Age of North Africa, PLOS ONE, vol.13, issue.10, p.e0202021, 2018.

C. K. Brain, The Hunters or the Hunted?: An Introduction to African Cave Taphonomy, 1981.

C. K. Brain and P. Shipman, The Swartkrans bone tools. Swartkrans. A Cave's Chronicle of Early Man, Transvaal Museum, pp.195-215, 1993.

T. G. Bromage, Interpretation of scanning electron microscopic images of abraded forming bone surfaces, American Journal of Physical Anthropology, vol.64, issue.2, pp.161-178, 1984.

E. Brühl, Lower Palaeolithic Small Tools in Europe and the Levant, British Archaeological Reports, pp.49-64, 2003.

H. T. Bunn, E. M. Kroll, S. H. Ambrose, A. K. Behrensmeyer, L. R. Binford et al., Systematic Butchery by Plio/Pleistocene Hominids at Olduvai Gorge, Tanzania [and Comments and Reply], Current Anthropology, vol.27, issue.5, pp.431-452, 1986.

S. D. Capaldo and R. J. Blumenschine, A Quantitative Diagnosis of Notches Made by Hammerstone Percussion and Carnivore Gnawing on Bovid Long Bones, American Antiquity, vol.59, issue.4, pp.724-748, 1994.

F. Errico, LA VIE SOCIALE DE L'ART MOBILIER PALÉOLITHIQUE. MANIPULATION, TRANSPORT, SUSPENSION DES OBJETS ON OS, BOIS DE CERVIDÉS, IVOIRE, Oxford Journal of Archaeology, vol.12, issue.2, pp.145-174, 1993.

F. D'errico and L. Backwell, Assessing the function of early hominin bone tools, Journal of Archaeological Science, vol.36, issue.8, pp.1764-1773, 2009.
URL : https://hal.archives-ouvertes.fr/halshs-00453450

F. D'errico and L. R. Backwell, Possible evidence of bone tool shaping by Swartkrans early hominids, Journal of Archaeological Science, vol.30, issue.12, pp.1559-1576, 2003.
URL : https://hal.archives-ouvertes.fr/halshs-00445516

F. Errico, L. R. Backwell, and L. R. Berger, Bone tool use in termite foraging by early hominids and its impact on our understanding of early hominid behaviour. South Afr, J. Sci, vol.97, pp.71-75, 2001.

F. D'errico, L. R. Backwell, and L. Wadley, Identifying regional variability in Middle Stone Age bone technology: The case of Sibudu Cave, Journal of Archaeological Science, vol.39, issue.7, pp.2479-2495, 2012.
URL : https://hal.archives-ouvertes.fr/hal-01842831

R. A. Dart, The Bone Tool-Manufacturing Ability of Australopithecus Prometheus, American Anthropologist, vol.62, issue.1, pp.134-143, 1960.

I. De-la-torre, The Early Stone Age lithic assemblages of Gadeb (Ethiopia) and the Developed Oldowan/early Acheulean in East Africa, Journal of Human Evolution, vol.60, issue.6, pp.768-812, 2011.

I. De-la-torre, R. M. Albert, A. Arroyo, R. Macphail, L. J. Mchenry et al., New excavations at the HWK EE site: Archaeology, paleoenvironment and site formation processes during late Oldowan times at Olduvai Gorge, Tanzania, Journal of Human Evolution, vol.120, pp.140-202, 2018.

I. De-la-torre and R. Mora, Technological behaviour in the early Acheulean of EF-HR (Olduvai Gorge, Tanzania), Journal of Human Evolution, vol.120, pp.329-377, 2018.

I. De-la-torre and R. Mora, Oldowan technological behaviour at HWK EE (Olduvai Gorge, Tanzania), Journal of Human Evolution, vol.120, pp.236-273, 2018.

A. L. Deino, 40Ar/39Ar dating of Bed I, Olduvai Gorge, Tanzania, and the chronology of early Pleistocene climate change, Journal of Human Evolution, vol.63, issue.2, pp.251-273, 2012.

A. L. Deino, C. Heil, J. King, L. J. Mchenry, I. G. Stanistreet et al., Chronostratigraphy and age modeling of Pleistocene drill cores from the Olduvai Basin, Tanzania (Olduvai Gorge Coring Project), Palaeogeography, Palaeoclimatology, Palaeoecology, p.109990, 2020.

V. Dobosi, G. Cavarretta, P. Gioia, M. Mussi, and . Palombo, Ex proboscideis-proboscidean remains as raw material at four Palaeolithic sites, Hungary, The world of Elephants. Proceedings of the First International Congress. Consiglio Nazionale Delle Ricerche, pp.429-431, 2001.

A. Echassoux, Comportements de subsistance et modifications osseuses à l?aube de l?Acheuléen à Konso, Éthiopie, L'Anthropologie, vol.116, issue.3, pp.291-320, 2012.

M. Gümrükçü and M. C. Pante, Assessing the effects of fluvial abrasion on bone surface modifications using high-resolution 3-D scanning, Journal of Archaeological Science: Reports, vol.21, pp.208-221, 2018.

R. L. Hay, Geology of the Olduvai Gorge: A Study of Sedimentation in a Semiarid Basin, 1976.

G. Haynes, Mammoths, mastodonts, and elephants: Biology, behavior, and the fossil record. Gary Haynes, 1991, Cambridge University Press, Cambridge, xi + 413 pp., $69.95 (clothbound), $27.95 (paperbound), Geoarchaeology, vol.9, issue.5, pp.419-419, 1994.

C. S. Henshilwood, F. D'errico, C. W. Marean, R. G. Milo, and R. Yates, An early bone tool industry from the Middle Stone Age at Blombos Cave, South Africa: implications for the origins of modern human behaviour, symbolism and language, Journal of Human Evolution, vol.41, issue.6, pp.631-678, 2001.

A. I. Herries, J. M. Martin, A. B. Leece, J. W. Adams, G. Boschian et al., Contemporaneity of Australopithecus, Paranthropus, and early Homo erectus in South Africa, Science, vol.368, issue.6486, p.eaaw7293, 2020.

W. Huang, Z. Zhang, R. Fu, B. Chen, J. Liu et al., Bone artifacts and ornaments from Xiaogushan site of Haicheng, Acta Anthropol. Sin, vol.5, pp.259-266, 1986.

M. Inizan, M. Reduron-ballinger, and H. Roche, Technology and Terminology of Knapped Stone, CREP, 1999.

C. C. Johnson, J. K. Njau, D. Van-damme, K. Schick, and N. Toth, PALEOECOLOGIC SIGNIFICANCE OF MALACOFAUNA, OLDUVAI GORGE, TANZANIA, PALAIOS, vol.31, issue.6, pp.319-326, 2016.

R. G. Klein, Anthropology: What do we know about neanderthals and Cro-Magnon man?, Am. Scholar, vol.52, pp.386-392, 1983.

R. G. Klein, The Human Career: Human Biological and Cultural Origins, 1999.

K. Kuman and R. J. Clarke, Stratigraphy, artefact industries and hominid associations for Sterkfontein, Member 5, Journal of Human Evolution, vol.38, issue.6, pp.827-847, 2000.

M. Leakey and D. A. Roe, Olduvai Gorge, Excavations in Beds III, IV and the Masek Beds, vol.5, 1994.

M. D. Leakey, Olduvai Gorge, Excavations in Beds I and II, vol.3, 1960.

M. Leakey, Modified bones from Beds III and IV, Excavations in Beds III, IV and the Masek Beds, vol.5, pp.311-314, 1994.

D. Mania and U. Mania, Bilzingsleben Homo erectus, his culture and his environment. The most important results of research, Lower Palaeolithic Small Tools in Europe and the Levant, pp.29-48, 2003.

S. Mcbrearty and A. S. Brooks, The revolution that wasn't: a new interpretation of the origin of modern human behavior, J. Hum. Evol, vol.39, pp.453-563, 2000.

H. M. Mchenry, C. C. Brown, and L. J. Mchenry, Fossil hominin ulnae and the forelimb of Paranthropus, Am. J. Phys. Anthropol, vol.134, pp.209-218, 2007.

L. J. Mchenry and I. G. Stanistreet, Tephrochronology of Bed II, Olduvai Gorge, Tanzania, and placement of the Oldowan-Acheulean transition, J. Hum. Evol, vol.120, pp.7-18, 2018.

A. Moigne, P. Valensi, P. Auguste, J. García-solano, A. Tuffreau et al., Bone retouchers from Lower Palaeolithic sites: Terra Amata, Orgnac 3, Cagny-l'Epinette and Cueva del Angel, Quaternary International, vol.409, pp.195-212, 2016.
URL : https://hal.archives-ouvertes.fr/hal-02100004

J. K. Njau and R. J. Blumenschine, A diagnosis of crocodile feeding traces on larger mammal bone, with fossil examples from the Plio-Pleistocene Olduvai Basin, Tanzania, J. Hum. Evol, vol.50, pp.142-162, 2006.

J. K. Njau and R. J. Blumenschine, Crocodylian and mammalian carnivore feeding traces on hominid fossils from FLK 22 and FLK NN 3, Plio-Pleistocene, Olduvai Gorge, Tanzania, J. Hum. Evol, vol.63, pp.408-417, 2012.

J. K. Njau, E. W. Herrmann, L. Ruck, M. Pante, P. Farrugia et al., Core stratigraphy constrains Bed IV archaeological record at HEB site, Olduvai Gorge, Tanzania. Palaeogeogr. Palaeoclimatol. Palaeoecol, vol.552, p.109773, 2020.

M. C. Pante, The larger mammal fossil assemblage from JK2, Bed III, Olduvai Gorge, Tanzania: implications for the feeding behavior of Homo erectus, Journal of Human Evolution, vol.64, issue.1, pp.68-82, 2013.

M. C. Pante, The larger mammal fossil assemblage from JK2, Bed III, Olduvai Gorge, Tanzania: implications for the feeding behavior of Homo erectus, J. Hum. Evol, vol.64, pp.68-82, 2013.

M. C. Pante and R. J. Blumenschine, Fluvial transport of bovid long bones fragmented by the feeding activities of hominins and carnivores, J. Archaeol. Sci, vol.37, pp.846-854, 2010.

M. C. Pante and I. De-la-torre, A hidden treasure of the Lower Pleistocene at Olduvai Gorge, Tanzania: the Leakey HWK EE assemblage, J. Hum. Evol, vol.120, pp.114-139, 2018.

M. C. Pante, J. K. Njau, B. Hensley-marschand, T. L. Keevil, C. Martín-ramos et al., The carnivorous feeding behavior of early Homo at HWK EE, Bed II, Olduvai Gorge, Tanzania, J. Hum. Evol, vol.120, pp.215-235, 2018.

K. A. Prassack, M. C. Pante, J. K. Njau, and I. De-la-torre, The paleoecology of Pleistocene birds from Middle Bed II, at Olduvai Gorge, Tanzania, and the environmental context of the Oldowan-Acheulean transition, J. Hum. Evol, vol.120, pp.32-47, 2018.

J. Pétillon, What are these barbs for? Preliminary study on the function of the Upper Magdalenian barbed weapon tips, P@lethnology, vol.1, pp.66-97, 2009.

T. Proffitt, Is there a Developed Oldowan A at Olduvai Gorge? A diachronic analysis of the Oldowan in Bed I and Lower-Middle Bed II at Olduvai Gorge, Tanzania, J. Hum. Evol, vol.120, pp.92-113, 2018.

A. M. Radmilli and G. Boschian, Gli Scavi a Castel di Guido: Il Più Antico Glacimento di Cacciatori del Paleolitico Inferire nell'Agro Romano. Istituto Italiano di Preistoria e Protostoria, 1996.

F. Rivals, K. T. Uno, F. Bibi, M. C. Pante, J. Njau et al., Dietary traits of the ungulates from the HWK EE site at Olduvai Gorge (Tanzania): Diachronic changes and seasonality, J. Hum. Evol, vol.120, pp.203-214, 2018.

R. Rabinovich, O. Ackermann, E. Aladjem, R. Barkai, R. Biton et al., Elephants at the Middle Pleistocene Acheulian open-air site of Revadim Quarry, Israel. Quat. Int, pp.183-197, 2012.

M. B. Roberts and S. A. Parfitt, Boxgrove: A Middle Pleistocene Hominid Site at Eartham Quarry, 1999.

J. Rosell, R. Blasco, G. Campeny, J. C. Díez, R. A. Alcalde et al., Bone as a technological raw material at the Gran Dolina site, J. Hum. Evol, vol.61, pp.125-131, 2011.

J. Rosell, R. Blasco, J. Fernández-peris, E. Carbonell, R. Barkai et al., Recycling bones in the Middle Pleistocene: Some reflections from Gran Dolina TD10-1 (Spain), Bolomor Cave (Spain) and Qesem Cave (Israel), Quat. Int, vol.361, pp.297-312, 2015.

K. Sano, Y. Beyene, S. Katoh, D. Koyabu, H. Endo et al., A 1.4-million-year-old bone handaxe from Konso, Ethiopia, shows advanced tool technology in the early Acheulean, Proc. Natl. Acad. Sci, 2020.
URL : https://hal.archives-ouvertes.fr/jpa-00218514

P. Shipman, Altered bones from Olduvai Gorge, Tanzania: Techniques, problems, and implications of their recognition, Bone Modification. Center for the Study of the First Americans, pp.247-258, 1989.

P. Shipman and J. J. Rose, Bone tools: an experimental approach, Scanning Electron Microscopy in Archaeology. British Archaeological Reports, pp.303-335, 1988.

R. C. Stammers, M. V. Caruana, and A. I. Herries, The first bone tools from Kromdraai and stone tools from Drimolen, and the place of bone tools in the South African Earlier Stone Age, Quat. Int, vol.495, pp.87-101, 2018.

I. G. Stanistreet, Fine resolution of early hominin time, Beds I and II, Olduvai Gorge, Tanzania, J. Hum. Evol, vol.63, pp.300-308, 2012.

I. G. Stanistreet, L. J. Mchenry, H. Stollhofen, and I. De-la-torre, Bed II Sequence Stratigraphic context of EF-HR and HWK EE archaeological sites, and the Oldowan/Acheulean succession at Olduvai Gorge, Tanzania, J. Hum. Evol, vol.120, pp.19-31, 2018.

I. G. Stanistreet, H. Stollhofen, A. L. Deino, L. J. Mchenry, N. P. Toth et al., New Olduvai Basin stratigraphy and stratigraphic concepts revealed by OGCP cores into the Palaeolake Olduvai depocentre, Tanzania, Palaeogeography, Palaeoclimatology, Palaeoecology, vol.554, p.109751, 2020.

C. Szmidt, J. Pétillon, P. Cattelain, C. Normand, and C. Schwab, Premiéres dates radiocarbone pour le Magdalénien d'Isturitz (Pyrénées Atlantiques), Bull. la Société Préhistorique Francaise, vol.106, pp.588-592, 2009.

N. Tisnerat-laborde, H. Valladas, and E. Ladier, Nouvelles datations carbone 14 en SMA pour le Magdalénien supérieur de la vallée de l, Aveyron. Préhist. Ariégeoise, vol.52, pp.129-136, 1997.

V. Tourloukis, N. Thompson, E. Panagopoulou, D. Giusti, G. E. Konidaris et al., Lithic artifacts and bone tools from the Lower Palaeolithic site Marathousa 1, Quat. Int, vol.497, pp.47-64, 2018.

K. T. Uno, F. Rivals, F. Bibi, M. Pante, J. Njau et al., Large mammal diets and paleoecology across the Oldowan-Acheulean transition at Olduvai Gorge, Tanzania from stable isotope and tooth wear analyses, J. Hum. Evol, vol.120, pp.76-91, 2018.

T. Van-kolfschoten, S. A. Parfitt, J. Serangeli, and S. M. Bello, Lower Paleolithic bone tools from the 'Spear Horizon' at Schöningen (Germany), J. Hum. Evol, vol.89, pp.226-263, 2015.

P. Villa and L. Bartram, Flaked bone from a hyena den, vol.8, pp.143-159, 1996.

P. Villa and F. Errico, Bone and ivory points in the Lower and Middle Paleolithic of Europe, J. Hum. Evol, vol.41, pp.69-112, 2001.
URL : https://hal.archives-ouvertes.fr/halshs-00444085

J. E. Yellen, Barbed bone points: tradition and continuity in Saharan and Sub-Saharan Africa, Afr. Archaeol. Rev, vol.15, pp.173-198, 1998.

J. E. Yellen, A. S. Brooks, E. Cornelissen, M. J. Mehlman, and K. Stewart, A Middle Stone Age worked bone industry from Katanda, Upper Semliki Valley, Zaire. Science, vol.268, pp.553-556, 1995.

S. Zhang, F. Errico, L. R. Backwell, Y. Zhang, F. Chen et al., Ma'anshan cave and the origin of bone tool technology in China, J. Archaeol. Sci, vol.65, pp.57-69, 2016.

Y. Zhang, M. C. Stiner, R. Dennell, C. Wang, S. Zhang et al., Zooarchaeological perspectives on the Chinese Early and Late Paleolithic from the Ma'anshan site, J. Archaeol. Sci, vol.37, pp.2066-2077, 2010.