P. L. Ames, The morphology of the syrinx in passerine birds, vol.37, 1971.

C. H. Greenwalt, . Bird, and . Song, Acoustics and Physiology

R. M. Casey and A. S. Gaunt, Theoretical models of the avian syrinx, J Theor Biol, 1985.

D. N. Düring, A. Ziegler, C. K. Thompson, A. Ziegler, C. Faber et al., The songbird syrinx morphome: A three-dimensional, high-resolution, interactive morphological map of the zebra finch vocal organ, BMC Biol, vol.11, p.1, 2013.

A. S. Gaunt, An hypothesis concerning the relationship of syringeal structure to vocal abilities, Auk, vol.100, pp.853-62, 1983.

A. S. King, Functional anatomy of the syrinx, pp.105-92, 1989.

O. N. Larsen and F. Goller, Direct observation of syringeal muscle function in songbirds and a parrot, J Exp Biol, 2002.

T. Riede and F. Goller, Peripheral mechanisms for vocal production in birds -differences and similarities to human speech and singing, vol.115, pp.69-80, 2011.

C. Elemans, J. H. Rasmussen, C. T. Herbst, D. N. Düring, S. A. Zollinger et al., Universal mechanisms of sound production and control in birds and mammals, Nat Commun, vol.6, 2015.

T. H. Huxley, A manual of the anatomy of vertebrated animals. D. Appleton and company

I. R. Titze, The physics of small-amplitude oscillation of the vocal folds, J Acoust Soc Am, vol.83, issue.4, pp.1536-52, 1988.

J. Van-den-berg, Myoelastic-Aerodynamic Theory of Voice Production, J Speech Lang Hear Res, vol.1, issue.3, p.227, 1958.

B. H. Story, The Handbook of Speech Production, 2015.

C. Elemans, I. Spierts, U. K. Müller, J. L. Van-leeuwen, and F. Goller, Superfast muscles control dove's trill, Nature, 2004.

C. Elemans, A. F. Mead, L. C. Rome, and F. Goller, Superfast vocal muscles control song production in songbirds, PLoS One, 2008.

C. Elemans, The singer and the song: The neuromechanics of avian sound production, Current Opinion in Neurobiology, vol.28, pp.172-180, 2014.

D. N. Düring, B. J. Knörlein, and C. Elemans, In situ vocal fold properties and pitch prediction by dynamic actuation of the songbird syrinx, Sci Rep, 2017.

T. Riede and F. Goller, Peripheral mechanisms for vocal production in birds -differences and similarities to human speech and singing, Brain Lang, vol.115, issue.1, pp.69-80, 2010.

M. Miskimen, Sound Production in Passerinc Birds, Auk, vol.68, pp.493-504, 1951.

O. N. Larsen and F. Goller, Role of syringeal vibrations in bird vocalizations, Proc R Soc B Biol Sci, 1999.

F. Goller and O. N. Larsen, A new mechanism of sound generation in songbirds, Proc Natl Acad Sci, vol.94, pp.14787-91, 1997.

D. N. Düring and C. Elemans, Embodied Motor Control of Avian Vocal Production, pp.119-57, 2016.

V. R. Ohms, P. C. Snelderwaard, C. Ten-cate, and G. Beckers, Vocal tract articulation in zebra finches, PLoS One, vol.5, issue.7, 2010.

G. Beckers, B. S. Nelson, and R. A. Suthers, Vocal-Tract Filtering by Lingual Articulation in a Parrot, Curr Biol, vol.14, pp.1592-1599, 2004.

W. J. Hoese, J. Podos, N. C. Boetticher, and S. Nowicki, Vocal tract function in birdsong production: experimental manipulation of beak movements, J Exp Biol, vol.203, pp.1845-55, 2000.

R. A. Suthers, How birds sing and why it matters, Nature's Music Sci Birdsong, pp.272-95, 2004.

R. A. Suthers, Peripheral vocal mechanisms in birds: Are songbirds special?, Netherlands J Zool, vol.51, issue.2, pp.217-259, 2001.

R. A. Suthers, Contributions to birdsong from the left and right sides of the intact syrinx, Nature, vol.347, pp.473-480, 1990.

R. A. Suthers and S. A. Zollinger, From brain to song: The vocal organ and vocal tract, Neurosci Birdsong, pp.78-98, 2008.

F. Goller and R. A. Suthers, Role of syringeal muscles in gating airflow and sound production in singing brown thrashers, J Neurophysiol, 1996.

B. Prince, T. Riede, and F. Goller, Sexual dimorphism and bilateral asymmetry of syrinx and vocal tract in the European starling (Sturnus vulgaris), J Morphol, 2011.

T. Riede and F. Goller, Functional morphology of the sound-generating labia in the syrinx of two songbird species, J Anat, vol.216, pp.23-36, 2010.

B. Miller and D. B. Miller, Two-voice phenomenon in Birds: Further evidence, Auk, vol.94, pp.567-72, 1977.

A. H. Krakauer, M. Tyrrell, K. Lehmann, N. Losin, F. Goller et al., Vocal and anatomical evidence for two-voiced sound production in the greater sage-grouse Centrocercus urophasianus, J Exp Biol, 2009.

L. Favaro, M. Gamba, C. Alfieri, D. Pessani, and A. G. Mcelligott, Vocal individuality cues in the African penguin (Spheniscus demersus): A source-filter theory approach, Sci Rep, 2015.

L. Favaro, M. Gamba, C. Gili, and D. Pessani, Acoustic correlates of body size and individual identity in banded penguins, PLoS One, 2017.

M. Gamba, O. Friard, and C. Giacoma, Vocal Tract Morphology Determines Species-Specific Features in Vocal Signals of Lemurs (Eulemur), Int J Primatol, 2012.

T. Aubin, P. Jouventin, and C. Hildebrand, Penguins use the two-voice system to recognize each other, Proc Biol Sci, vol.267, pp.1081-1088, 2000.
URL : https://hal.archives-ouvertes.fr/hal-02555352

P. Jouventin, Visual and vocal signals in penguins, their evolution and adaptive characters, Fortschr Verhaltensforsch, vol.24, p.148, 1982.

H. J. Kriesell, T. Aubin, V. Planas-bielsa, M. Benoiste, F. Bonadonna et al., Sex identification in King Penguins Aptenodytes patagonicus through morphological and acoustic cues, Ibis, 1859.
URL : https://hal.archives-ouvertes.fr/hal-01721443

T. Lengagne, J. Lauga, and T. Aubin, Intra-syllabic acoustic signatures used by the king penguin in parent-chick recognition: an experimental approach, J Exp Biol, vol.204, pp.663-72, 2001.
URL : https://hal.archives-ouvertes.fr/hal-02555326

T. Aubin and P. Jouventin, How to vocally identify kin in a crowd: The penguin model, 2002.

P. Robisson, Vocalizations in aptenodytes penguins : Application of the two-voice theory, Auk, vol.109, issue.3, pp.654-662, 1992.

I. Wilden, H. Herzel, G. Peters, and G. Tembrock, Subharmonics, biphonation, and deterministic chaos in Mammal vocalization, Bioac, vol.9, issue.3, pp.171-96, 1988.

J. F. Meckel, Theilung der Luftröhre durch eine Scheidewand bei der Fettgans (Aptenodytes demersa), pp.81-75

M. Watson, The Voyage of H.M.S. Challenger. In: 1st ed. London: Longmans and Co, p.1885

P. M. Zeek, Double trachea in penguins and sea lions, Anat Rec, vol.111, issue.3, pp.327-370, 1951.

J. Davenport, T. T. Jones, T. M. Work, and G. H. Balazs, Unique characteristics of the trachea of the juvenile leatherback turtle facilitate feeding, diving and endothermy, J Exp Mar Bio Ecol, 2014.

D. G. Homberger, The avian tongue and larynx: Multiple functions in nutrition and vocalisation, Proceedings of the 22nd International Ornithological Congress, pp.94-113, 1999.

C. Casteleyn, P. Cornillie, S. Van-cruchten, W. Van-den-broeck, C. Van-ginneken et al., Anatomy of the upper respiratory tract in domestic birds, with emphasis on vocalization, Journal of Veterinary Medicine Series C, 2018.

K. Kobayashi, M. Kumakura, and K. Yoshimura, Fine Structure of the Tongue and Lingual Papillae of the Penguin, vol.61, 1998.

T. Lengagne, T. Aubin, J. Lauga, and P. Jouventin, How do king penguins (Aptenodytes patagonicus) apply the mathematical theory of information to communicate in windy conditions?, Proc R Soc B Biol Sci, 1999.
URL : https://hal.archives-ouvertes.fr/hal-00173757

T. Lengagne, P. Jouventin, and T. Aubin, Finding One 's Mate In A King Penguin Colony: Efficiency of Acoustic Communication, Behaviour, vol.136, pp.833-879, 1999.

W. T. Fitch, Acoustic exaggeration of size in birds via tracheal elongation: Comparative and theoretical analyses, J Zool, 1999.

C. Elemans, M. Muller, O. N. Larsen, and J. L. Van-leeuwen, Amplitude and frequency modulation control of sound production in a mechanical model of the avian syrinx, J Exp Biol, 2009.

I. R. Titze, Iowa City: National Center for Voice and Speech, vol.409, 2000.

T. Riede and F. Goller, Morphological basis for the evolution of acoustic diversity in oscine songbirds, Proc R Soc B Biol Sci, 2014.

R. A. Suthers and F. Goller, Motor correlates of vocal diversity in songbirds, Curr Ornithol, vol.14, pp.235-88, 1997.

S. A. Zollinger, T. Riede, and . Suthers-r-a, Two-voice complexity from a single side of the syrinx in northern mockingbird Mimus polyglottos vocalizations, J Exp Biol, vol.211, pp.1978-91, 2008.

F. Goller and T. Riede, Integrative physiology of fundamental frequency control in birds, Journal of Physiology, vol.107, pp.230-272, 2013.

C. Elemans, R. Zaccarelli, and H. Herzel, Biomechanics and control of vocalization in a nonsongbird, J R Soc Interface, vol.5, issue.24, pp.691-703, 2008.

M. Budka and T. S. Osiejuk, Formant Frequencies are Acoustic Cues to Caller Discrimination and are a Weak Indicator of the Body Size of Corncrake Males, Ethology, 2013.

P. J. Ponganis, S. Leger, J. Scadeng, and M. , Penguin lungs and air sacs: implications for baroprotection, oxygen stores and buoyancy, J Exp Biol, 2015.

B. Cozzi, P. Bagnoli, F. Acocella, and M. L. Costantino, Structure and biomechanical properties of the trachea of the striped dolphin Stenella coeruleoalba: Evidence for evolutionary adaptations to diving, Anat Rec -Part A Discov Mol Cell Evol Biol, 2005.

C. Moore, M. Moore, S. Trumble, M. Niemeyer, B. Lentell et al., A comparative analysis of marine mammal tracheas, J Exp Biol, 2014.

A. G. Hocken, Post-mortem examination of penguins, Dep Conserv New Zealand, Sci Intern Ser, vol.65, pp.1-25, 2002.

S. L. Cook and T. G. Forrest, Sounds produced by nesting leatherback sea turtles (Dermochelys coriacea), Herpetol Rev, vol.36, issue.4, pp.387-90, 2005.

N. Mrosovsky, Spectrographs of the Sounds of Leatherback Turtles, Herpetologica, vol.28, issue.3, pp.256-264, 1972.

P. K. Anderson and R. Barclay, Acoustic Signals of Solitary Dugongs: Physical Characteristics and Behavioral Correlates, J Mammal, vol.76, issue.4, pp.1226-1263, 1995.

K. Pütz and Y. Cherel, The diving behaviour of brooding king penguins (Aptenodytes patagonicus) from the Falkland Islands: Variation in dive profiles and synchronous underwater swimming provide new insights into their foraging strategies, Mar Biol, 2005.

R. P. Wilson, A. Steinfurth, Y. Ropert-coudert, A. Kato, and M. Kurita, Lip-reading in remote subjects: An attempt to quantify and separate ingestion, breathing and vocalisation in free-living animals using penguins as a model, Mar Biol, vol.140, issue.1, pp.17-27, 2002.

B. D. Metscher, MicroCT for developmental biology: A versatile tool for high-contrast 3D imaging at histological resolutions, Dev Dyn, 2009.

B. D. Metscher, Micro CT for comparative morphology: Simple staining methods allow highcontrast 3D imaging of diverse non-mineralized animal tissues, BMC Physiol, 2009.

B. Ruthensteiner and M. Heß, Embedding 3D models of biological specimens in PDF publications, Microsc Res Tech, 2008.

K. Delord, C. Barbraud, and H. Weimerskirch, Long-term trends in the population size of king penguins at Crozet archipelago: Environmental variability and density dependence?, Polar Biol, vol.27, issue.12, pp.793-800, 2004.
URL : https://hal.archives-ouvertes.fr/hal-00186860

T. Lengagne, T. Aubin, P. Jouventin, and J. Lauga, Acoustic communication in a king penguin colony: Importance of bird location within the colony and of the body position of the listener, Polar Biol, 1999.

T. Lengagne, T. Aubin, P. Jouventin, and J. Lauga, Perceptual salience of individually distinctive features in the calls of adult king penguins, J Acoust Soc Am, 2000.
URL : https://hal.archives-ouvertes.fr/hal-02555373

R. Specht and . Avisoft-saslab-pro, Avisoft Bioacoustics, 2004.