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E. L. , R. S. , and W. M. , gratefully acknowledges support from an Investigator Award from the Gordon and Betty Moore Foundation Marine Microbiology Initiative (GBMF3783) and from a grant from the Simons Foundation (542395) as part of the Principles of Microbial Ecosystems (PriME) Collaborative. W.M.D. gratefully acknowledges a startup grant from the University of Sheffield's Imagine: Imaging Life initiative and a New Investigator Grant from the Biotechnology and Biological Sciences Research Council (BB/ R018383/1)

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