C. H. Albert, B. Rayfield, M. Dumitru, and A. Gonzalez, Applying network theory to prioritize multispecies habitat networks that are robust to climate and land-use change, Conservation Biology, vol.31, pp.1383-1396, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01681596

G. Alligand, S. Hubert, T. Legendre, F. Millard, A. Müller et al., THEMA Évaluation environnementale -Étude d'aide à la définition des mesures ERC, %C3 %A9ma %20-%20Guide %20d %E2 %80 %99aide %20 %C3 %A0 %20la %20d %C3 %A9finition %20des %20mesures %20ERC.pdf, 2018.

J. Amsallem, R. Sordello, L. Billon, and V. Sylvie, Bilan des Schémas régionaux de cohérence écologique en France : quels apports méthodologiques pour l'identification et la cartographie de la Trame verte et bleue ?, Sciences Eaux & Territoires, vol.25, pp.4-11, 2018.

F. Ascensão, F. Mestre, and A. M. Barbosa, Prioritizing road defragmentation using graphbased tools, Landscape and Urban Planning, vol.192, 2019.

C. Avon and L. Bergès, Outils d'analyse de la connectivité des habitats, Convention DEB-MEDDE / Irstea, 2013.

C. Avon and L. Bergès, Prioritization of habitat patches for landscape connectivity conservation differs between least-cost and resistance distances, Landscape Ecology, vol.31, pp.1551-1565, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01393253

C. Avon, L. Bergès, and P. Roche, Comment analyser la connectivité écologique des trames vertes ? Cas d'étude en région méditerranéenne, Sciences Eaux & Territoires, vol.14, pp.14-19, 2014.

M. Belisle, Measuring landscape connectivity: The challenge of behavioral landscape ecology, Ecology, vol.86, issue.8, pp.1988-1995, 2005.

A. Bergsten and A. Zetterberg, To model the landscape as a network : A practitioner's perspective, Landscape and Urban Planning, vol.119, pp.35-43, 2013.

, Intégrer la connectivité paysagère dans la séquence ERC : une approche par la

, VertigO -la revue électronique en sciences de l'environnement, vol.19, p.2019

L. Bezombes, S. Gaucherand, T. Spiegelberger, V. Gouraud, and C. Kerbiriou, A set of organized indicators to conciliate scientific knowledge, offset policies requirements and operational constraints in the context of biodiversity offsets, Ecological Indicators, vol.93, pp.1244-1252, 2018.

C. Bigard, S. Pioch, and J. D. Thompson, The inclusion of biodiversity in environmental impact assessment : Policy-related progress limited by gaps and semantic confusion, Journal of Environmental Management, pp.35-45, 0200.

L. Boitani, A. Falcucci, L. Maiorano, and C. Rondinini, Ecological networks as conceptual frameworks or operational tools in conservation, Conservation Biology, vol.21, pp.1414-1422, 2007.

S. Briggs and M. D. Hudson, Determination of significance in Ecological Impact Assessment : Past change, current practice and future improvements, Environmental Impact Assessment Review, vol.38, pp.16-25, 2013.

D. J. Bruggeman, M. L. Jones, F. Lupi, and K. T. Scribner, Landscape equivalency analysis : methodology for estimating spatially explicit biodiversity credits, Environmental Management, vol.36, issue.4, pp.518-534, 2005.

J. W. Bull, A. Gordon, J. E. Watson, and M. Maron, Seeking convergence on the key concepts in « no net loss' policy, Journal of Applied Ecology, vol.53, pp.1686-1693, 2016.

C. Gamarra, M. J. , J. P. Lassoie, and J. Milder, Accounting for no net loss : A critical assessment of biodiversity offsetting metrics and methods, Journal of Environmental Management, vol.220, pp.36-43, 2018.

, Convention on Biological Diversity (CBD), 2010, Decision UNEP/CBD/COP/DEC/X/2 Adopted by the Conference of the Parties to the Convention on Biological Diversity at Its Tenth Meeting, 2018.

C. Clauzel and C. Bonnevalle, Apports de la modélisation spatiale pour la gestion de la trame verte et bleue, Cybergeo : European Journal of Geography, 2019.

C. Clauzel, D. Xiqing, W. Gongsheng, P. Giraudoux, and L. Li, Assessing the impact of road developments on connectivity across multiple scales : Application to Yunnan snub-nosed monkey conservation, Biological Conservation, vol.192, pp.207-217, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01207624

A. Coulon, J. Aben, S. C. Palmer, V. M. Stevens, T. Callens et al., A stochastic movement simulator improves estimates of landscape connectivity, vol.96, pp.2203-2213, 2015.

K. R. Crooks and M. Sanjayan, Connectivity conservation, Conservation Biology, vol.712, 2006.

T. Dalang and A. M. Hersperger, Trading connectivity improvement for area loss in patchbased biodiversity reserve networks, Biological Conservation, vol.148, pp.116-125, 2012.

O. Darses, C. Cantuarias-villessuzanne, ;. Cgdd, and P. Mtes, Sites naturels de compensation, un outil prometteur au service de la biodiversité, 2017.

B. De-la-fuente, M. C. Mateo-sánchez, G. Rodríguez, A. Gastón, R. Pérez-de-ayala et al., Natura 2000 sites, public forests and riparian corridors : The connectivity backbone of forest green infrastructure, Land Use Policy, vol.75, pp.429-441, 2018.

, VertigO -la revue électronique en sciences de l'environnement, vol.19, p.2019

H. Dehouck and J. Ansallem, Analyse des méthodes de précision des continuités écologiques à l'échelle locale en France. Irstea -UMR TETIS, Centre de ressources Trame verte et bleue. 96p, 2018.

R. Duflot, C. Avon, P. Roche, and L. Bergès, Combining habitat suitability models and spatial graphs for more effective landscape conservation planning : an applied methodological framework and a species case study, Journal for Nature Conservation, vol.46, pp.38-47, 2018.

, Landscapes in transition. An account of 25 years of land cover change in Europe, Publications Office of the European Union, vol.88, 2017.

F. Eigenbrod, S. J. Hecnar, and L. Fahrig, Accessible habitat: an improved measure of the effects of habitat loss and roads on wildlife populations, Landscape Ecology, vol.23, issue.2, pp.159-168, 2008.

W. F. Fagan, J. M. Calabrese-;-k, M. Crooks, and . Sanjayan, Quantifying connectivity : balancing metric performance with data requirements, Connectivity Conservation, pp.297-317, 2006.

L. Fahrig, Ecological responses to habitat fragmentation per se, Evolution, and Systematics, vol.48, pp.1-23, 2017.

J. C. Foltête, C. Clauzel, and G. Vuidel, A software tool dedicated to the modelling of landscape networks, Environmental Modelling & Software, vol.38, pp.316-327, 2012.

J. C. Foltête, X. Girardet, and C. Clauzel, A methodological framework for the use of landscape graphs in land-use planning, Landscape and Urban Planning, vol.124, pp.140-150, 2014.

J. Foltête, How ecological networks could benefit from landscape graphs : A response to the paper by Spartaco Gippoliti and Corrado Battisti, Land Use Policy, vol.80, pp.391-394, 2019.

R. T. Forman, Land mosaics : the ecology of landscapes and regions, vol.632, 1995.

P. Gibbons and D. B. Lindenmayer, Offsets for land clearing : No net loss or the tail wagging the dog ?, Ecological Management & Restoration, vol.8, issue.1, pp.26-31, 2007.

X. Girardet, Paysage & infrastructures de transport -modélisation des impacts des infrastructures sur les réseaux écologiques, vol.261, 2014.

M. S. Gurrutxaga and . Saura, Prioritizing highway defragmentation locations for restoring landscape connectivity, Environmental Conservation, vol.41, issue.2, pp.157-164, 2014.

, Summary for policymakers of the global assessment report on biodiversity and ecosystem services of the Intergovernmental Science-Policy Platform on Biodiversity and Ecosystem Services, IPBES secretariat, Intergovernmental Science-Policy Platform on Biodiversity and Ecosystem Services (IPBES), 2019.

C. Jacob, F. Quétier, J. Aronson, S. Pioch, and H. Levrel, Vers une politique française de compensation des impacts sur la biodiversité plus efficace : défis et perspectives VertigO -la revue électronique en sciences de l'environnement, vol.14, 2014.

R. H. Jongman, M. Külvik, and I. Kristiansen, Landscape and Urban Planning, vol.68, issue.2, pp.305-319, 2004.

, Intégrer la connectivité paysagère dans la séquence ERC : une approche par la

, VertigO -la revue électronique en sciences de l'environnement, vol.19, p.2019

J. M. Kiesecker, H. Copeland, A. Pocewicz, and B. Mckenney, Development by design : blending landscape-level planning with the mitigation hierarchy, Frontiers in Ecology and the Environment, vol.8, issue.5, pp.261-266, 2010.

J. M. Kiesecker, H. Copeland, A. Pocewicz, N. Nibbelink, B. Mckenney et al.,

. Stroud, A framework for implementing biodiversity offsets : selecting sites and determining scale, Bioscience, vol.59, pp.77-84, 2009.

H. Kujala, A. L. Whitehead, W. K. Morris, and B. A. Wintle, Towards strategic offsetting of biodiversity loss using spatial prioritization concepts and tools : A case study on mining impacts in Australia, Biological Conservation, vol.192, pp.513-521, 2015.

A. Laita, J. S. Kotiaho, and M. Monkkonen, Graph-theoretic connectivity measures: what do they tell us about connectivity?, Landscape Ecology, vol.26, pp.951-967, 2011.

S. Lapoint, P. Gallery, M. Wikelski, and R. Kays, Animal behavior, cost-based corridor models, and real corridors, Landscape Ecology, vol.28, pp.1615-1630, 2013.

T. G. Martin, M. A. Burgman, F. Fidler, P. M. Kuhnert, S. Low-choy et al., Eliciting expert knowledge in conservation science, Conservation Biology, vol.26, pp.29-38, 2012.

M. Maron, S. Brownlie, J. W. Bull, M. C. Evans, A. Hase et al.,

G. , The many meanings of no net loss in environmental policy, Nature Sustainability, vol.1, issue.1, pp.19-27, 2018.

B. H. Mcrae, B. G. Dickson, T. H. Keitt, and V. B. Shah, Using circuit theory to model connectivity in ecology, evolution, and conservation, Ecology, vol.89, pp.2712-2724, 2008.

M. Meurant, A. Gonzalez, A. Doxa, and C. H. Albert, Selecting surrogate species for connectivity conservation, Biological Conservation, vol.227, pp.326-334, 2018.
URL : https://hal.archives-ouvertes.fr/hal-01914756

. Ministère-de-l'écologie, Lignes directrices nationales sur la séquence éviter, réduire et compenser les impacts sur les milieux naturels, ReferenceS, CGDD DEB, vol.232, 2013.

A. Mimet, C. Clauzel, and J. C. Foltête, Locating wildlife crossings for multispecies connectivity across linear infrastructures, Landscape Ecology, vol.31, pp.1955-1973, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01297307

T. Newbold, L. N. Hudson, A. P. Arnell, S. Contu, A. Palma et al.,

. Purvis, Has land use pushed terrestrial biodiversity beyond the planetary boundary ? À global assessment, Science, vol.353, pp.288-291, 2016.

S. Pradon, M. Mereau, A. Baghli, D. Labarraque, and P. Thievent, OPTIFLUX : une innovation en matière d'ingénierie des infrastructures de transports face aux enjeux de la biodiversité, vol.877, pp.90-93, 2010.

F. Quétier, B. Regnery, and H. Levrel, No net loss of biodiversity or paper offsets ? A critical review of the French no net loss policy, Environmental Science & Policy, vol.38, pp.120-131, 2014.

B. Rayfield, D. Pelletier, M. Dumitru, J. A. Cardille, and A. Gonzalez, Multipurpose habitat networks for short-range and long-range connectivity : a new method combining graph and circuit connectivity, Methods in Ecology and Evolution, vol.7, issue.2, pp.222-231, 2016.

B. Rayfield, M. J. Fortin, and A. Fall, The sensitivity of least-cost habitat graphs to relative cost surface values, Landscape Ecology, vol.25, pp.519-532, 2010.

, Intégrer la connectivité paysagère dans la séquence ERC : une approche par la

, VertigO -la revue électronique en sciences de l'environnement, vol.19, p.2019

B. Rayfield, M. J. Fortin, and A. Fall, Connectivity for conservation : a framework to classify network measures, Ecology, vol.92, pp.847-858, 2011.

B. Regnery, F. Quétier, N. Cozannet, S. Gaucherand, A. Laroche et al.,

. Kerbiriou, Mesures compensatoires pour la biodiversité : comment améliorer les dossiers environnementaux et la gouvernance ?, Sciences Eaux et Territoires, vol.12, pp.1-8, 2013.

Y. Sahraoui, J. C. Foltête, and C. Clauzel, A multi-species approach for assessing the impact of land-cover changes on landscape connectivity, Landscape Ecology, vol.32, pp.1819-1835, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01577903

L. Santini, S. Saura, and C. Rondinini, A composite network approach for assessing multispecies connectivity: an application to road defragmentation prioritisation, Plos One, vol.11, p.10, 2016.

S. Saura, Node self-connections in network metrics, Ecology Letters, vol.21, issue.2, pp.319-320, 2018.

S. Saura, B. De-la-fuente, ;. S. Gergel, and M. G. Turner, Connectivity as the amount of reachable habitat : conservation priorities and the roles of habitat patches in landscape networks, pp.229-254, 2017.

S. Saura, C. Estreguil, C. Mouton, and M. Rodriguez-freire, Network analysis to assess landscape connectivity trends : Application to European forests, Ecological Indicators, vol.11, issue.2, pp.407-416, 1990.

S. Saura and J. Torné, Conefor Sensinode 2.2 : A software package for quantifying the importance of habitat patches for landscape connectivity, Environmental Modelling & Software, vol.24, pp.135-139, 2009.

S. Saura and L. , A new habitat availability index to integrate connectivity in landscape conservation planning : Comparison with existing indices and application to a case study, Landscape and Urban Planning, vol.83, pp.91-103, 2007.

S. Saura and L. Rubio, A common currency for the different ways in which patches and links can contribute to habitat availability and connectivity in the landscape, Ecography, vol.33, pp.523-537, 2010.

S. C. Sawyer, C. W. Epps, and J. S. Brashares, Placing linkages among fragmented habitats: do least-cost models reflect how animals use landscapes, Journal of Applied Ecology, vol.48, pp.668-678, 2011.

L. R. Tambosi, A. C. Martensen, M. C. Ribeiro, and J. P. Metzger, A framework to optimize biodiversity restoration efforts based on habitat amount and landscape connectivity, Restoration Ecology, vol.22, issue.2, pp.169-177, 2014.

S. Tarabon, L. Bergès, T. Dutoit, and F. Isselin-nondedeu, Environmental impact assessment of development projects improved by merging species distribution and habitat connectivity modelling, Journal of Environmental Management, vol.241, pp.439-449, 2109.
URL : https://hal.archives-ouvertes.fr/hal-02136980

S. Tarabon, L. Bergès, T. Dutoit, and F. Isselin-nondedeu, Maximizing habitat connectivity in the mitigation hierarchy. A case study on three terrestrial mammals in an urban environment, Journal of Environmental Management, vol.243, pp.340-349, 2019.
URL : https://hal.archives-ouvertes.fr/hal-02136967

P. D. Taylor, L. Fahrig, K. Henein, and G. Merriam, Connectivity is a vital element of landscape structure, Oikos, vol.68, pp.571-573, 1993.

J. G. Underwood, Combining landscape-level conservation planning and biodiversity offset programs : a case study, Environmental Management, vol.47, pp.121-129, 2011.

D. L. Urban, E. S. Minor, E. A. Treml, and R. S. Schick, Graph models of habitat mosaics, Ecology Letters, vol.12, pp.260-273, 2009.

, Intégrer la connectivité paysagère dans la séquence ERC : une approche par la

, VertigO -la revue électronique en sciences de l'environnement, vol.19, p.2019

V. Vasas, T. Magura, F. Jordan, and B. Tothmeresz, Graph theory in action: evaluating planned highway tracks based on connectivity measures, Landscape Ecology, vol.24, pp.581-586, 2009.

J. Wawrzyczek, R. Lindsay, M. J. Metzger, and F. Quétier, The ecosystem approach in ecological impact assessment: lessons learned from windfarm developments on peatlands in Scotland, Environmental Impact Assessment Review, vol.72, pp.157-165, 2018.

W. Wende, G. Tucker, F. Quétier, M. Rayment, and M. Darbi, Biodiversity Offsets-The European Perspective on No Net Loss of Biodiversity and Ecosystem Services, vol.252, 2018.

K. A. Zeller, K. Mcgarigal, and A. R. Whiteley, Estimating landscape resistance to movement : a review, Landscape Ecology, vol.27, pp.777-797, 2012.

K. A. Zeller, M. K. Jennings, T. W. Vickers, H. B. Ernest, S. A. Cushman et al., Are all data types and connectivity models created equal? Validating common connectivity approaches with dispersal data, Diversity and Distributions, vol.24, pp.868-879, 2018.

, La séquence ERC a pour principe d'évaluer l'ensemble des impacts écologiques d'un aménagement, mais les critères et indicateurs permettant de quantifier les impacts sur la connectivité paysagère ne sont pas satisfaisants. Nous proposons une démarche opérationnelle pour évaluer les impacts environnementaux d'un aménagement sur la connectivité des habitats à l'échelle du paysage. Cette démarche est basée sur la théorie des graphes et l'indice EC de « quantité d'habitat atteignable » pour les espèces, qui tient compte de la quantité d'habitat, de la configuration spatiale des taches d'habitats et des capacités de dispersion des espèces. Les trois principaux enjeux de la séquence ERC sont redéfinis au regard de cette démarche, les politiques environnementales dans différents pays mettent en oeuvre la séquence Eviter-Réduire-Compenser

, Nous discutons des atouts et des limites de cette approche pour la séquence ERC et fournissons Intégrer la connectivité paysagère dans la séquence ERC : une approche par la

, VertigO -la revue électronique en sciences de l'environnement, vol.19

U. R. Irstea and . Lessem, Université Grenoble Alpes, 2, rue de la papeterie, courriel : laurent.berges@irstea.fr CATHERINE AVON Semperfloris, vol.76

U. R. Irstea and . Lessem, Université Grenoble Alpes, 2, rue de la papeterie, BP, vol.76

. Ladyss, U. Cnrs, and P. De, courriel : celine.clauzel@univ-paris-diderot.fr Intégrer la connectivité paysagère dans la séquence ERC : une approche par la, vol.75013

, VertigO -la revue électronique en sciences de l'environnement, P.O. Box, vol.19, issue.2, p.40014, 2019.

. Théma, UMR 6049 CNRS, vol.32, p.25030

F. Besançon,

U. R. Irstea and . Lessem, Université Grenoble Alpes, 2, rue de la papeterie, BP, vol.76

. Théma, UMR 6049 CNRS, vol.32, p.25030

F. Besançon,

U. R. Irstea and . Lessem, thomas.spiegelberger@irstea.fr Intégrer la connectivité paysagère dans la séquence ERC : une approche par la, BP, vol.76

, VertigO -la revue électronique en sciences de l'environnement, vol.19, p.2019