G. T. Baker and R. L. Sprott, Biomarkers of aging. Experimental Gerontology, vol.23, pp.223-323, 1988.

G. Barja, The flux of free radical attack through mitochondrial DNA is related to aging 325 rate, Aging (Milano), vol.12, pp.342-355, 2000.

G. Barja and A. Herrero, Oxidative damage to mitochondrial DNA is inversely related to 327 maximum life span in the heart and brain of mammals, FASEB J, vol.14, pp.312-318, 2000.

D. Bates, M. Mächler, B. Bolker, and S. Walker, Fitting linear mixed-effects models 329 using lme4, 2014.

R. Bonduriansky, A. Maklakov, F. Zajitschek, and R. Brooks, Sexual selection, sexual 331 conflict and the evolution of ageing and life span, Functional Ecology, vol.22, pp.443-453, 2008.

J. Campisi and F. Di-fagagna, Cellular senescence: when bad things happen to 333 good cells, Nature reviews. Molecular cell biology, vol.8, p.729, 2007.

Y. Cao, H. Li, S. Deb, and J. Liu, TERT regulates cell survival independent of 335 telomerase enzymatic activity, Oncogene, vol.21, pp.3130-3138, 2002.

M. A. Chebbi, T. Becking, B. Moumen, I. Giraud, C. Gilbert et al., The 337 Genome of Armadillidium vulgare (Crustacea, Isopoda) Provides Insights into Sex 338 Chromosome Evolution in the Context of Cytoplasmic Sex Determination, Molecular 339 Biology and Evolution, 2019.

F. Chevalier, J. Herbinière-gaboreau, J. Bertaux, M. Raimond, F. Morel et al., The Immune Cellular Effectors of Terrestrial Isopod Armadillidium vulgare: 342 Meeting with Their Invaders, Wolbachia. PLOS ONE, vol.6, p.18531, 2011.

L. Cheynel, J. Lemaître, J. Gaillard, B. Rey, G. Bourgoin et al., , 2017.

, Immunosenescence patterns differ between populations but not between sexes in a 345 long-lived mammal, Scientific Reports, vol.7

C. Chiu and C. B. Harley, Replicative Senescence and Cell Immortality: The Role of 347 Telomeres and Telomerase, Experimental Biology and Medicine, vol.214, pp.99-106, 1997.

B. B. De-jesus and M. A. Blasco, Assessing cell and organ senescence biomarkers, Circulation research, vol.349, pp.97-109, 2012.

C. Depeux, A. Samba-louaka, C. Braquart-varnier, J. Moreau, J. Lemaître et al., Impact of temperature and photoperiod impact on survival and biomarkers 352 of senescence in common woodlouse, bioRxiv, 2019.

G. P. Dimri, X. Lee, G. Basile, M. Acosta, G. Scott et al., A 354 biomarker that identifies senescent human cells in culture and in aging skin in vivo, PNAS, vol.355, pp.9363-9367, 1995.

T. Finkel and N. J. Holbrook, Oxidants, oxidative stress and the biology of ageing, Nature, vol.357, pp.239-247, 2000.

R. K. Gary and S. M. Kindell, Quantitative assay of senescence-associated ? -359 galactosidase activity in mammalian cell extracts, Analytical biochemistry, vol.343, pp.329-360, 2005.

S. W. Geiser, Further observations on the sex-ratios of terrestrial isopods, Field Lab, vol.3, pp.362-369, 1934.

N. M. Gomes, J. W. Shay, and W. E. Wright, Telomere biology in Metazoa, FEBS 364 Letters, vol.584, pp.3741-3751, 2010.

D. Harman, Aging: a theory based on free radical and radiation chemistry, Journal of 366 gerontology, vol.11, pp.298-300, 1956.

U. Herbig, M. Ferreira, L. Condel, D. Carey, and J. M. Sedivy, Cellular senescence in 368 aging primates, Science, vol.311, p.1257, 2006.

Y. Hsieh and C. Hsu, Honeybee trophocytes and fat cells as target cells for cellular 370 senescence studies, Experimental Gerontology, vol.46, pp.233-240, 2011.

K. Itahana, J. Campisi, and G. P. Dimri, Methods to detect biomarkers of cellular 372 senescence: the senescence-associated ? -galactosidase assay, Biological Aging: 373 Methods and Protocols, pp.21-31, 2007.

A. R. Ives and . Robert, For testing the significance of regression coefficients, 375 go ahead and log transform count data, Methods in Ecology and Evolution, vol.6, pp.828-376, 2015.

C. Lawless, C. Wang, D. Jurk, A. Merz, T. Zglinicki et al., Quantitative 378 assessment of markers for cell senescence, Exp. Gerontol, vol.45, pp.772-778, 2010.

L. Liu, S. M. Bailey, M. Okuka, P. Muñoz, C. Li et al., Telomere 380 lengthening early in development, Nature Cell Biology, vol.9, pp.1436-1441, 2007.

G. A. Marais, J. Gaillard, C. Vieira, I. Plotton, D. Sanlaville et al., Sex gap in aging and longevity: can sex chromosomes play a role? Biology of 383 Sex Differences 9, 2018.

. Cc-by-nc-nd, International license It is made available under a (which was not peer-reviewed) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. The copyright holder for this preprint, 2019.

G. Morgan, Telomerase regulation and the intimate relationship with aging, 2013.

O. H. Paris and F. A. Pitelka, Population characteristics of the terrestrial isopod 387 Armadillidium vulgare in California grassland, Ecology, vol.43, pp.229-248, 1962.

Y. Park, Y. Kim, and D. Stanley, Cellular immunosenescence in adult male crickets, p.389, 2011.

, Gryllus assimilis. Archives of Insect Biochemistry and Physiology, vol.76, pp.185-194

J. D. Podlevsky, C. J. Bley, R. V. Omana, X. Qi, J. J. Chen et al., The Telomerase 391 Database, Nucleic Acids Research, vol.36, pp.339-343, 2007.

R. C. Team, R: A language and environment for statistical computing. Vienna: R 393 Foundation for Statistical Computing, 2014.

J. L. Ram and A. J. Costa, Invertebrates as Model Organisms for Research on Aging 395 Biology * . In: Conn's Handbook of Models for Human Aging, pp.445-452, 2018.

H. M. Robertson and K. H. Gordon, Canonical TTAGG-repeat telomeres and telomerase 397 in the honey bee, Apis mellifera, Genome Research, vol.16, pp.1345-1351, 2006.

J. Rodriguez, V. Boulo, E. Mialhe, and E. Bachere, Characterisation of shrimp 399 haemocytes and plasma components by monoclonal antibodies, Journal of Cell, vol.400, pp.1043-1050, 1995.

C. Schumpert, J. Nelson, E. Kim, J. L. Dudycha, and R. C. Patel, Telomerase Activity 402 and Telomere Length in Daphnia, vol.10, p.127196, 2015.

J. W. Shay and W. E. Wright, Senescence and immortalization: role of telomeres and 404 telomerase, Carcinogenesis, vol.26, pp.867-874, 2005.

K. W. Shertzer and S. P. Ellner, State-dependent energy allocation in variable 406 environments: life history evolution of a rotifer, Ecology, vol.83, pp.2181-2193, 2002.

M. Sicard, F. Chevalier, M. D. Vlechouver, D. Bouchon, P. Grève et al., Variations of immune parameters in terrestrial isopods: a matter of gender, 409 aging and Wolbachia, Naturwissenschaften, vol.97, pp.819-826, 2010.

A. Simm, N. Nass, B. Bartling, B. Hofmann, R. Silber et al., 411 Potential biomarkers of ageing, Biological Chemistry, vol.389, 2008.

R. L. Sprott, Biomarkers of aging and disease: Introduction and definitions, 2010.

, Experimental Gerontology, vol.45, pp.2-4

D. Stanley, Aging and immunosenescence in invertebrates, Invertebrate Survival 415 Journal, vol.9, pp.102-109, 2012.

C. G. Steel, Mechanisms of coordination between moulting and reproduction in 417 terrestrial isopod crustacea, The Biological Bulletin, vol.159, pp.206-218, 1980.

M. Tidière, J. Gaillard, D. W. Müller, L. B. Lackey, O. Gimenez et al., Does sexual selection shape sex differences in longevity and senescence 420 patterns across vertebrates? A review and new insights from captive ruminants: 421 SEXUAL SELECTION AND SENESCENCE, Evolution, vol.69, pp.3123-3140, 2015.

A. E. Vinogradov, Male Reproductive Strategy and Decreased Longevity, Acta 423 Biotheoretica, vol.46, pp.157-160, 1998.

C. Wang, D. Jurk, M. Maddick, G. Nelson, C. Martin-ruiz et al., Aging Cell 8: was not peer-reviewed) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. The copyright holder for this preprint, 2009.