S. Ahmed, J. Cock, and E. Pessia, A Haploid System of Sex Determination in the Brown Alga Ectocarpus sp., Current Biology, vol.24, issue.17, pp.1945-1957, 2014.
DOI : 10.1016/j.cub.2014.07.042

URL : https://hal.archives-ouvertes.fr/hal-01132642

L. Anderson and J. Snider, Cytological and genetic barriers in mosses, Journal of the Hattori Botanical Laboratory, vol.52, pp.241-254, 1982.

E. Anderson and E. Thompson, A model-based method for identifying species hybrids using multilocus genetic data, Genetics, vol.160, pp.1217-1229, 2002.

S. Arnaud-haond and K. Belkhir, genclone: a computer program to analyse genotypic data, test for clonality and describe spatial clonal organization, Molecular Ecology Notes, vol.106, issue.1, pp.15-17, 2007.
DOI : 10.1111/j.1471-8286.2006.01522.x

N. Arnheim, M. Krystal, and R. Schmickel, Molecular evidence for genetic exchanges among ribosomal genes on nonhomologous chromosomes in man and apes., Proceedings of the National Academy of Sciences, pp.7323-7327, 1980.
DOI : 10.1073/pnas.77.12.7323

N. Barton and G. Hewitt, Analysis of Hybrid Zones, Annual Review of Ecology and Systematics, vol.16, issue.1, pp.113-148, 1985.
DOI : 10.1146/annurev.es.16.110185.000553

K. Belkhir, P. Borsa, L. Chikhi, N. Raufaste, and F. Bonhomme, GENETIX 4.05, logiciel sous Windows TM pour la génétique des populations, Laboratoire Génome, 2004.

L. Bergström, A. Tatarenkov, K. Johannesson, R. Jönsson, and L. Kautsky, GENETIC AND MORPHOLOGICAL IDENTIFICATION OF FUCUS RADICANS SP. NOV. (FUCALES, PHAEOPHYCEAE) IN THE BRACKISH BALTIC SEA1., Journal of Phycology, vol.12, issue.5, pp.1025-1038, 2005.
DOI : 10.1111/j.1529-8817.2004.04085.x

N. Bierne, P. Borsa, and C. Daguin, Introgression patterns in the mosaic hybrid zone between Mytilus edulis and M. galloprovincialis, Molecular Ecology, vol.112, issue.2, pp.447-461, 2003.
DOI : 10.1007/s002270050342

E. Billard, E. Serrão, and G. Pearson, (Fucaceae, Phaeophyceae) and their putative hybrids, European Journal of Phycology, vol.15, issue.4, pp.397-407, 2005.
DOI : 10.1046/j.1365-2540.2001.00968.x

URL : http://sapientia.ualg.pt/bitstream/10400.1/4156/1/Analysis%20of%20sexual%20phenotype%20and%20prezygotic%20fertility%20in%20natural%20populations%20of%20Fucus%20spiralis%2c%20F.%20vesiculosus%20%28Fucaceae%2c%20Phaeophyceae%29%20and%20their%20putative%20hybrids.pdf

J. Cock, L. Sterck, and P. Rouzé, The Ectocarpus genome and the independent evolution of multicellularity in brown algae, Nature, vol.10, issue.7298, pp.617-621, 2010.
DOI : 10.2216/i0031-8884-39-3-258.1

URL : https://hal.archives-ouvertes.fr/cea-00906990

J. Cock, O. Godfroy, N. Macaisne, A. Peters, and S. Coelho, Evolution and regulation of complex life cycles: a brown algal perspective, Current Opinion in Plant Biology, vol.17, pp.1-6, 2014.
DOI : 10.1016/j.pbi.2013.09.004

S. Coelho, A. Peters, and B. Charrier, Complex life cycles of multicellular eukaryotes: New approaches based on the use of model organisms, Gene, vol.406, issue.1-2, pp.152-170, 2007.
DOI : 10.1016/j.gene.2007.07.025

J. Coyer, G. Smith, and R. Andersen, EVOLUTION OF MACROCYSTIS SPP. (PHAEOPHYCEAE) AS DETERMINED BY ITS1 AND ITS2 SEQUENCES1,, Journal of Phycology, vol.37, issue.4, pp.574-585, 2001.
DOI : 10.1046/j.1529-8817.2001.037001574.x

J. Coyer, A. Peters, G. Hoarau, W. Stam, and J. Olsen, Hybridization of the marine seaweeds, Fucus serratus and Fucus evanescens (Heterokontophyta: Phaeophyceae) in a 100-year-old zone of secondary contact, Proceedings of the Royal Society B: Biological Sciences, vol.269, issue.1502, pp.1829-1834, 2002.
DOI : 10.1098/rspb.2002.2093

J. Coyer, G. Hoarau, M. Oudot-le-secq, W. Stam, and J. Olsen, A mtDNA-based phylogeny of the brown algal genus Fucus (Heterokontophyta; Phaeophyta), Molecular Phylogenetics and Evolution, vol.39, issue.1, pp.209-222, 2006.
DOI : 10.1016/j.ympev.2006.01.019

J. Coyer, G. Hoarau, W. Stam, and J. Olsen, Hybridization and introgression in a mixed population of the intertidal seaweeds Fucus evanescens and F. serratus, Journal of Evolutionary Biology, vol.13, issue.6, pp.2322-2333, 2007.
DOI : 10.1038/nrg1269

J. Coyer, G. Hoarau, and J. Costa, Evolution and diversification within the intertidal brown macroalgae Fucus spiralis/F. vesiculosus species complex in the North Atlantic, Molecular Phylogenetics and Evolution, vol.58, issue.2, pp.283-296, 2011.
DOI : 10.1016/j.ympev.2010.11.015

C. Destombe, M. Valero, and M. Guillemin, Delineation of Two Sibling Red Algal Species, Gracilaria Gracilis and Gracilaria Dura (Gracilariales, Rhodophyta), Using Multiple DNA Markers: Resurrection of the Species G. Dura Previously Described in the Northern Atlantic 200???Years Ago1, Journal of Phycology, vol.38, issue.4, pp.720-727, 2010.
DOI : 10.2216/i0031-8884-34-2-113.1

M. Dorken and C. Eckert, Severely reduced sexual reproduction in northern populations of a clonal plant, Decodonverticillatus (Lythraceae), Journal of Ecology, vol.85, issue.3, pp.339-350, 2001.
DOI : 10.2307/2960610

G. Dover, Concerted evolution, molecular drive and natural selection, Current Biology, vol.4, issue.12, pp.1165-1166, 1994.
DOI : 10.1016/S0960-9822(00)00265-7

C. Engel, C. Daguin, and E. Serrão, Genetic entities and mating system in hermaphroditic Fucus spiralis and its close dioecious relative F. vesiculosus (Fucaceae, Phaeophyceae), Molecular Ecology, vol.40, issue.261, pp.2033-2046, 2005.
DOI : 10.1111/j.0014-3820.2003.tb01494.x

G. Evanno, S. Regnaut, and J. Goudet, Detecting the number of clusters of individuals using the software structure: a simulation study, Molecular Ecology, vol.16, issue.8, pp.2611-2620, 2005.
DOI : 10.1098/rspb.2001.1600

P. Famá, J. Olsen, W. Stam, and G. Procaccini, High levels of intra- and inter-individual polymorphism in the rDNA ITS1 of Caulerpa racemosa (Chlorophyta), European Journal of Phycology, vol.35, issue.4, pp.349-356, 2000.
DOI : 10.1017/S0967026200002948

A. Geoffroy, S. Mauger, D. Jode, A. , L. Gall et al., (Ectocarpales, Phaeophyceae) along the Brittany coast, Journal of Phycology, vol.6, issue.3, pp.480-489, 2015.
DOI : 10.1371/journal.pone.0019402

URL : https://hal.archives-ouvertes.fr/hal-01147191

T. Giraud, G. Refrégier, L. Gac, M. De-vienne, D. Hood et al., Speciation in fungi, Fungal Genetics and Biology, vol.45, issue.6, pp.791-802, 2008.
DOI : 10.1016/j.fgb.2008.02.001

G. Hoarau, J. Coyer, M. Giesbers, A. Jüterbock, and J. Olsen, Pre-zygotic isolation in the macroalgal genus Fucus from four contact zones spanning 100-10 000 years: a tale of reinforcement?, Royal Society Open Science, vol.70, issue.2, p.140538, 2015.
DOI : 10.3354/meps143211

H. Ito, T. Langenhorst, R. Ogden, and M. Inoue-murayama, Population genetic diversity and hybrid detection in captive zebras, Scientific Reports, vol.6, issue.1, p.13171, 2015.
DOI : 10.1111/j.1471-8286.2006.01433.x

URL : http://doi.org/10.1038/srep13171

M. Jakobsson and N. Rosenberg, CLUMPP: a cluster matching and permutation program for dealing with label switching and multimodality in analysis of population structure, Bioinformatics, vol.23, issue.14, pp.1801-1806, 2007.
DOI : 10.1093/bioinformatics/btm233

K. Johannesson, Inverting the null-hypothesis of speciation: a marine snail perspective, Evolutionary Ecology, vol.123, issue.1, pp.5-16, 2009.
DOI : 10.1111/j.0014-3820.2004.tb00869.x

T. Jombart, adegenet: a R package for the multivariate analysis of genetic markers, Bioinformatics, vol.24, issue.11, pp.1403-1405, 2008.
DOI : 10.1093/bioinformatics/btn129

URL : https://hal.archives-ouvertes.fr/hal-00428105

M. Knight, XV.???Studies in the Ectocarpace??. II. The Life-history and Cytology of Ectocarpus siliculosus, Dillw, Transactions of the Royal Society of Edinburgh, vol.x, issue.02, pp.307-332, 1930.
DOI : 10.1017/S0080456800013223

L. Kohn, Mechanisms of Fungal Speciation, Annual Review of Phytopathology, vol.43, issue.1, pp.279-308, 2005.
DOI : 10.1146/annurev.phyto.43.040204.135958

Y. Lammers, D. Kremer, and P. Brakefield, SNP genotyping for detecting the ???rare allele phenomenon??? in hybrid zones, Molecular Ecology Resources, vol.36, issue.2, pp.237-242, 2013.
DOI : 10.1111/j.1095-8312.1989.tb00495.x

P. Lavretsky, J. Peters, and K. Winker, Becoming pure: identifying generational classes of admixed individuals within lesser and greater scaup populations, Molecular Ecology, vol.17, issue.Suppl, pp.661-674, 2016.
DOI : 10.1111/j.1365-294X.2008.03737.x

C. Lexer, C. Buerkle, J. Joseph, B. Heinze, and M. Fay, Admixture in European Populus hybrid zones makes feasible the mapping of loci that contribute to reproductive isolation and trait differences, Heredity, vol.37, issue.2, pp.74-84, 2007.
DOI : 10.1126/science.244.4911.1490

P. Liao, H. Shih, and T. Yen, Molecular evaluation of interspecific hybrids between Acer albopurpurascens and A. buergerianum var. formosanum, Botanical Studies, vol.51, pp.413-420, 2010.

G. Liti, D. Barton, and E. Louis, Sequence Diversity, Reproductive Isolation and Species Concepts in Saccharomyces, Genetics, vol.174, issue.2, pp.839-850, 2006.
DOI : 10.1534/genetics.106.062166

URL : http://www.ncbi.nlm.nih.gov/pmc/articles/PMC1602076

K. Livingstone and L. Rieseberg, Chromosomal evolution and speciation: a recombination-based approach, New Phytologist, vol.75, issue.1, pp.107-112, 2004.
DOI : 10.1111/j.0014-3820.2003.tb01537.x

URL : http://www.ncbi.nlm.nih.gov/pmc/articles/PMC2601666

B. Mable and S. Otto, The evolution of life cycles with haploid and diploid phases, BioEssays, vol.92, issue.261, pp.453-462, 1998.
DOI : 10.1007/978-1-4684-1402-8_8

C. Maggs, H. Fletcher, and D. Fewer, Speciation in Red Algae: Members of the Ceramiales as Model Organisms, Integrative and Comparative Biology, vol.51, issue.3, pp.492-504, 2011.
DOI : 10.1093/icb/icr075

A. Montecinos, L. Couceiro, and A. Peters, in press. Species delimitation and phylogeographic analyses in the Ectocarpus subgroup siliculosi (Ectocarpales, Phaeophyceae), Journal of Phycology

C. Monteiro, E. Serrão, and G. Pearson, Prezygotic Barriers to Hybridization in Marine Broadcast Spawners: Reproductive Timing and Mating System Variation, PLoS ONE, vol.39, issue.2, p.35978, 2012.
DOI : 10.1371/journal.pone.0035978.g008

URL : http://doi.org/10.1371/journal.pone.0035978

D. Müller and G. Gassmann, Sexual hormone specificity in Ectocarpus and Laminaria (Phaeophyceae), Naturwissenschaften, vol.89, issue.9, pp.462-463, 1980.
DOI : 10.1007/BF00405647

D. Müller, Culture experiments on life cycle, nuclear phases, and sexuality of the brown alga Ectocarpus siliculosus, Planta, vol.24, issue.1, pp.39-54, 1967.
DOI : 10.1007/BF00380838

D. Müller, Studies on sexual compatibility betweenEctocarpus siliculosus (Phaeophyceae) from Chile and the Mediterranean Sea, Helgol??nder Meeresuntersuchungen, vol.14, issue.3-4, p.469, 1988.
DOI : 10.2216/i0031-8884-18-4-312.1

J. Neiva, G. Pearson, M. Valero, and E. Serrão, Surfing the wave on a borrowed board: range expansion and spread of introgressed organellar genomes in the seaweed Fucus ceranoides L., Molecular Ecology, vol.175, issue.21, pp.4812-4822, 2010.
DOI : 10.1093/oxfordjournals.molbev.a003747

E. Nielsen, L. Bach, and P. Kotlicki, hybridlab (version 1.0): a program for generating simulated hybrids from population samples, Molecular Ecology Notes, vol.15, issue.4, pp.971-973, 2006.
DOI : 10.1111/j.1471-8286.2006.01433.x

K. Niwa and A. Kobiyama, (Bangiales, Rhodophyta), Journal of Phycology, vol.8, issue.5, pp.897-900, 2014.
DOI : 10.1371/journal.pone.0073414

D. Paetkau, R. Slade, M. Burden, and A. Estoup, Genetic assignment methods for the direct, real-time estimation of migration rate: a simulation-based exploration of accuracy and power, Molecular Ecology, vol.16, issue.1, pp.55-65, 2004.
DOI : 10.1016/B978-012323445-2/50005-5

R. Peakall and P. Smouse, GenAlEx 6.5: genetic analysis in Excel. Population genetic software for teaching and research--an update, Bioinformatics, vol.28, issue.19, pp.2537-2539, 2012.
DOI : 10.1093/bioinformatics/bts460

R. Pereyra, L. Bergström, L. Kautsky, and K. Johannesson, Rapid speciation in a newly opened postglacial marine environment, the Baltic Sea, BMC Evolutionary Biology, vol.9, issue.1, p.70, 2009.
DOI : 10.1186/1471-2148-9-70

A. Peters, D. Marie, D. Scornet, B. Kloareg, M. Cock et al., (ECTOCARPALES, PHAEOPHYCEAE) AS A MODEL ORGANISM FOR BROWN ALGAL GENETICS AND GENOMICS, Journal of Phycology, vol.40, issue.6, pp.1079-1088, 2004.
DOI : 10.1111/j.1529-8817.2004.04058.x

A. Peters, D. Scornet, D. Müller, B. Kloareg, and J. Cock, (Phaeophyceae), European Journal of Phycology, vol.8, issue.3, pp.235-242, 2004.
DOI : 10.1073/pnas.242379899

URL : https://hal.archives-ouvertes.fr/hal-00925432

A. Peters, V. Wijk, S. Cho, and G. , Reinstatement of Ectocarpus crouaniorum Thuret in Le Jolis as a third common species of Ectocarpus (Ectocarpales, Phaeophyceae) in Western Europe, and its phenology at Roscoff, Brittany, Phycological Research, vol.29, issue.3, pp.157-170, 2010.
DOI : 10.1016/B978-0-12-372180-8.50042-1

A. Peters, A. Mann, and C. Cordova, Genetic diversity of Ectocarpus (Ectocarpales, Phaeophyceae) in Peru and northern Chile, the area of origin of the genome-sequenced strain, New Phytologist, vol.43, issue.1, pp.30-41, 2010.
DOI : 10.1080/09670260903383271

A. Pillmann, G. Woolcott, J. Olsen, W. Stam, and R. King, Inter- and intraspecific genetic variation in Caulerpa (Chlorophyta) based on nuclear rDNA ITS sequences, European Journal of Phycology, vol.32, issue.4, pp.379-386, 1997.
DOI : 10.1017/S0967026297001352

S. Piry and J. Cornuet, GENECLASS. A Program for Assignation and Exclusion Using Molecular Markers, URLB, 1999.

J. Pritchard, M. Stephens, and P. Donnelly, Inference of population structure using multilocus genotype data, Genetics, vol.155, pp.945-959, 2000.

M. Quillet, N. Madjidian, and Y. Griveau, Mapping genetic factors controlling pollen viability in an interspecific cross in Helianthus sect. Helianthus, Theoretical and Applied Genetics, vol.91, issue.8, pp.1195-1202, 1995.
DOI : 10.1007/BF00220929

B. Rannala and J. Mountain, Detecting immigration by using multilocus genotypes, Proceedings of the National Academy of Sciences, pp.9197-9201, 1997.
DOI : 10.1111/j.1469-1809.1995.tb01605.x

URL : https://www.ncbi.nlm.nih.gov/pmc/articles/PMC23111/pdf

L. Rieseberg, Chromosomal rearrangements and speciation, Trends in Ecology & Evolution, vol.16, issue.7, pp.351-358, 2001.
DOI : 10.1016/S0169-5347(01)02187-5

N. Rosenberg, distruct: a program for the graphical display of population structure, Molecular Ecology Notes, vol.298, issue.1, pp.137-138, 2004.
DOI : 10.1046/j.1471-8286.2003.00566.x

G. Russell, The ecology of some free-living Ectocarpaceae. Helgoländer wissenschaftliche Meeresuntersuchungen, pp.155-162, 1967.

G. Russell, The genus Ectocarpus in Britain II. The Free-living forms, Journal of the Marine Biological Association of the United Kingdom, vol.1, issue.01, pp.233-250, 1967.
DOI : 10.1002/iroh.19090020105

G. Russell, Formation of an ectocarpoid epiflora on blades of Laminaria digitata, Marine Ecology Progress Series, vol.11, pp.181-187, 1983.
DOI : 10.3354/meps011181

A. Savoie and G. Saunders, (Ceramiales, Rhodophyta) in the Northwest Atlantic, Molecular Ecology, vol.5, issue.23, pp.5927-5937, 2015.
DOI : 10.1006/mpev.1995.1027

M. Schilthuizen, R. Hoekstra, and E. Gittenberger, Selective increase of a rare haplotype in a land snail hybrid zone, Proceedings of the Royal Society B: Biological Sciences, vol.266, issue.1434, pp.2181-2185, 1999.
DOI : 10.1098/rspb.1999.0906

E. Serrão, L. Alice, and S. Brawley, EVOLUTION OF THE FUCACEAE (PHAEOPHYCEAE) INFERRED FROM nrDNA-ITS, Journal of Phycology, vol.35, issue.2, pp.382-394, 1999.
DOI : 10.1046/j.1529-8817.1999.3520382.x

J. Sites and C. Moritz, Chromosomal Evolution and Speciation Revisited, Systematic Zoology, vol.36, issue.2, pp.153-174, 1987.
DOI : 10.2307/2413266

P. Soltis and D. Soltis, The Role of Hybridization in Plant Speciation, Annual Review of Plant Biology, vol.60, issue.1, pp.561-588, 2009.
DOI : 10.1146/annurev.arplant.043008.092039

F. Spirito, D. Howard, and S. Berlocher, The role of chromosomal change in speciation In: Endless forms: Species and speciation, pp.320-329, 1998.

B. Stache, Sexual compatability and species concept in Ectocarpus siliculosus (Ectocarpales, Phaeophyceae) from Italy In: Evolutionary biogeography of the marine algae of, Garbary DJ, pp.173-186, 1990.

DOI : 10.1111/j.0022-3646.1997.00152.x

G. Stebbins, The Inviability, Weakness, and Sterility of Interspecific Hybrids, Advances in Genetics, vol.9, pp.147-215, 1958.
DOI : 10.1016/S0065-2660(08)60162-5

G. Stebbins, Chromosomal evolution in higher plants, 1971.

C. Turchetto, A. Segatto, J. Beduschi, S. Bonatto, and L. Freitas, Genetic differentiation and hybrid identification using microsatellite markers in closely related wild species, AoB Plants, vol.7, p.84, 2015.
DOI : 10.1093/aobpla/plv084

URL : http://doi.org/10.1093/aobpla/plv084

J. Vähä and C. Primmer, Efficiency of model-based Bayesian methods for detecting hybrid individuals under different hybridization scenarios and with different numbers of loci, Molecular Ecology, vol.38, issue.1, pp.63-72, 2006.
DOI : 10.1093/oxfordjournals.molbev.a003956

M. Valero, S. Richerd, V. Perrot, and C. Destombe, Evolution of alternation of haploid and diploid phases in life cycles, Trends in Ecology & Evolution, vol.7, issue.1, pp.25-29, 1992.
DOI : 10.1016/0169-5347(92)90195-H

W. Wagner, Some questions about natural hybrids in ferns, Botanica Helvetica, vol.97, pp.195-205, 1987.

A. Wallace, A. Klein, and A. Mathieson, (PHAEOPHYTA) IN A MAINE ESTUARY, Journal of Phycology, vol.15, issue.6, pp.1013-1027, 2004.
DOI : 10.1007/978-1-349-27910-4

B. Weir and C. Cockerham, Estimating F-statistics for the analysis of population structure. evolution, Evolution, pp.1358-1370, 1984.

G. Zardi, K. Nicastro, and F. Canovas, Adaptive Traits Are Maintained on Steep Selective Gradients despite Gene Flow and Hybridization in the Intertidal Zone, PLoS ONE, vol.191, issue.6, p.19402, 2011.
DOI : 10.1371/journal.pone.0019402.s001

G. Zuccarello, N. Schidlo, L. Mcivor, and M. Guiry, (Gigartinales, Rhodophyta) in Europe, European Journal of Phycology, vol.132, issue.4, pp.337-344, 2005.
DOI : 10.1515/BOT.2004.064