R. Alkemade, A. Wielemaker, S. A. De-jong, and A. J. Sandee, Experimental evidence for the role of bioturbation by the marine nematode Diplolaimella dievengatensis in stimulating the mineralization of Spartina anglica detritus, Marine Ecology Progress Series, vol.90, pp.149-155, 1992.
DOI : 10.3354/meps090149

B. Gapped and P. , a new generation of protein database search 534 programs, Nucleic Acids Research, vol.25, pp.3389-3402

G. F. Blanchard, S. Bouhet, B. Guarini, and J. , Properties of the dynamics of 541 intertidal microphytobentic biomass, Journal of the Marine Biological Association of the 542 UK, pp.1027-1028, 2002.

G. F. Blanchard, P. Sauriau, . Cariou-le, V. Gall, M. J. Garet et al., Kinetics of tidal resuspension of microbiota:testing the effects of sediment cohesiveness and bioturbation using flume experiments, Marine Ecology Progress Series, vol.151, pp.17-25, 1997.
DOI : 10.3354/meps151017

S. Borin, L. Brusetti, D. Daffonchio, E. Delaney, and F. Baldi, Biodiversity of prokaryotic 553 communities in sediments of different sub-basins of the Venice lagoon, pp.307-314, 2009.

J. P. Bowman and R. D. Mccuaig, Biodiversity, community structural shifts, and 556 biogeography of prokaryotes within Antarctic continental shelf sediment Applied and 557 Environmental Microbiology, pp.2463-2483, 2003.

J. F. Brouwer and L. J. De-stal, Daily fluctuations of exopolymers in cultures of the 559, 2002.

C. P. Brussaard, Optimization of Procedures for Counting Viruses by Flow Cytometry, Applied and Environmental Microbiology, vol.70, issue.3, pp.1506-1513, 2004.
DOI : 10.1128/AEM.70.3.1506-1513.2004

B. C. Cho and F. Azam, Major role of bacteria in biogeochemical fluxes in the ocean's interior, Nature, vol.332, issue.6163, pp.441-443, 1988.
DOI : 10.1038/332441a0

R. J. Chróst and B. Riemann, Storm-stimulated enzymatic decomposition of organic matter in benthic/pelagic coastal meso-cosms, Marine Ecology Progress Series, vol.108, pp.185-192, 1994.
DOI : 10.3354/meps108185

K. R. Clarke and R. W. Warwick, Change in Marine Communities: An Approach to Statistical 568 Analysis and Interpretation, 2001.

J. B. Cotner, T. H. Johengen, and B. A. Biddanda, Intense winter heterotrophic production stimulated by benthic resuspension, Limnology and Oceanography, vol.45, issue.7, pp.1672-1676, 2000.
DOI : 10.4319/lo.2000.45.7.1672

B. C. Crump, E. V. Armbrust, and J. A. Baross, Phylogenetic analysis of particle-attached 572, 1999.

R. Danovaro, A. Dell-'anno, A. Trucco, M. Serresi, and S. Vanucci, Determination of Virus Abundance in Marine Sediments, Applied and Environmental Microbiology, vol.67, issue.3, pp.1384-576, 2001.
DOI : 10.1128/AEM.67.3.1384-1387.2001

D. Brouwer, J. F. Stal, and L. J. , Short-term dynamics in microphytobenthos distribution 578 and associated extracellular carbohydrates in surface sediments of an intertidal mudflat, p.579, 2001.

M. Dubois, K. A. Gilles, J. K. Hamilton, P. A. Rebers, and F. Smith, Colorimetric Method for Determination of Sugars and Related Substances, Analytical Chemistry, vol.28, issue.3, pp.350-356, 1956.
DOI : 10.1021/ac60111a017

S. Duhamel and S. Jacquet, Flow cytometric analysis of bacteria- and virus-like particles in lake sediments, Journal of Microbiological Methods, vol.64, issue.3, pp.316-332, 2006.
DOI : 10.1016/j.mimet.2005.05.008

L. B. Fandino, L. Riemann, G. F. Steward, R. A. Long, and F. Azam, Variations in bacterial community structure during a dinoflagellate bloom analyzed by DGGE and 16S rDNA sequencing, Aquatic Microbial Ecology, vol.23, p.119, 2001.
DOI : 10.3354/ame023119

C. Fiordelmondo and A. Pusceddu, Short-term response of benthic bacteria and 591 nanoflagellates to sediment resuspension: an experimental study, Chemistry and Ecology, vol.592, issue.20, pp.107-121, 2004.

M. È. Garneau, W. F. Vincent, R. Terrado, and C. Lovejoy, Importance of particle- 594 associated bacterial heterotrophy in a coastal Arctic ecosystem, Journal of Marine Systems, vol.595, pp.75-185, 2009.

T. Garstecki, S. A. Wickham, and H. Arndt, Effects of Experimental Sediment Resuspension on a Coastal Planktonic Microbial Food Web, Estuarine, Coastal and Shelf Science, vol.55, issue.5, pp.751-598, 2002.
DOI : 10.1006/ecss.2001.0937

S. U. Gerbersdorf, R. Bittner, H. Lubarsky, W. Manz, and D. M. Paterson, Microbial 600 assemblages as ecosystem engineers of sediment stability, Journal of Soils and Sediments, vol.601, issue.9, pp.640-652, 2009.

J. F. Ghiglione, P. Conan, and M. Pujo?pay, Diversity of total and active free?living vs. 603 particle?attached bacteria in the euphotic zone of the NW Mediterranean Sea. FEMS 604 Microbiology Letters, pp.9-21, 2009.

N. Goto, O. Mitamura, and H. Terai, Biodegradation of photosynthetically produced 606 extracellular organic carbon from intertidal benthic algae, Journal of Experimental Marine Biology and Ecology, vol.607, issue.257, pp.73-86, 2001.

R. C. Grabowski, I. G. Droppo, and G. Wharton, Erodibility of cohesive sediment: The importance of sediment properties, Earth-Science Reviews, vol.105, issue.3-4, pp.101-120, 2011.
DOI : 10.1016/j.earscirev.2011.01.008

K. Guizien, F. Orvain, J. C. Duchene, L. Hir, and P. , Accounting for rough bed friction 611 factors of mud beds due to biological activity in erosion experiments, Journal of Hydraulic, vol.612, issue.138, pp.979-984, 2012.

K. Guizien, C. Dupuy, P. Ory, H. Montanié, H. Hartmann et al., Microorganism dynamics during a rising tide: Disentangling effects of resuspension and mixing with offshore waters above an intertidal mudflat, Journal of Marine Systems, vol.129, issue.129, pp.616-178
DOI : 10.1016/j.jmarsys.2013.05.010

URL : https://hal.archives-ouvertes.fr/hal-01248055

I. Hamels, K. Muylaert, G. Casteleyn, and W. Vyverman, Uncoupling of bacterial 618 production and flagellate grazing in aquatic sediments: a case study from an intertidal flat, p.619, 2001.

S. Hedtkamp, Shallow subtidal sand: Permeability, nutrient dynamics 623 microphytobenthos and organic matter. Christian-Albrechts-Universität, 2005.

M. Héral, C. Bacher, and J. M. Deslous-paoli, La Capacité trophique des bassins 625 ostréicoles (ed) L'homme et les ressources halieutiques : essai sur l'usage 626 d'une ressource renouvelable Ifremer, pp.225-259, 1989.

F. Inagaki, M. Suzuki, K. Takai, H. Oida, T. Sakamoto et al., Microbial Communities Associated with Geological Horizons in Coastal Subseafloor Sediments from the Sea of Okhotsk, Applied and Environmental Microbiology, vol.69, issue.12, pp.7224-630, 2003.
DOI : 10.1128/AEM.69.12.7224-7235.2003

B. S. Kim, B. K. Kim, J. H. Lee, M. Kim, Y. W. Lim et al., Rapid phylogenetic dissection of prokaryotic community structure in tidal flat using pyrosequencing, The Journal of Microbiology, vol.53, issue.4, pp.357-363, 2008.
DOI : 10.1007/s12275-008-0071-9

B. S. Kim, H. M. Oh, H. Kang, and J. Chun, Archaeal diversity in tidal flat sediment as 638 revealed by 16S rDNA analysis, Journal of Microbiology, vol.43, pp.144-151, 2005.

E. D. Kopczynski, M. M. Bateson, and D. M. Ward, Recognition of chimeric small-subunit 640, 1994.

A. Lapoussière, C. Michel, M. Starr, M. Gosselin, and M. Poulin, Role of free-living and particle-attached bacteria in the recycling and export of organic material in the Hudson Bay system, Journal of Marine Systems, vol.88, issue.3, pp.434-445, 2011.
DOI : 10.1016/j.jmarsys.2010.12.003

J. R. Lazier and K. H. Mann, Turbulence and the diffusive layers around small organisms, Deep Sea Research Part A. Oceanographic Research Papers, vol.36, issue.11, p.646, 1989.
DOI : 10.1016/0198-0149(89)90068-X

L. Hir, P. Cann, P. Jestin, H. Bassoulet, and P. , Instrumentation légère pour la mesure de 648 l " érodabilité des sédiments vaseux ou sablovaseux, Proc. IXémes Journées Nationales 649, 2006.

C. J. Lorenzen, A method for the continuous measurement of in vivo chlorophyll 654 concentration, Deep-Sea Research, vol.13, pp.223-227, 1966.

S. U. Gerbersdorf, The stabilisation potential of individual and mixed assemblages of 657 natural bacteria and microalgae, PloS one, vol.5, p.13794, 2010.

C. R. Woese, A new version of the RDP (ribosomal database project) Nucleic Acids 661 Research, pp.171-173, 1999.

C. Mallet and D. Debroas, Relations between organic matter and bacterial proteolytic 663 activity in sediment surface layers of a eutrophic lake, 1999.

C. Mallet and D. Debroas, Regulation of ?-and ?-glycolytic activities in the sediments of a 666 eutrophic lake, Microbial Ecology, vol.41, pp.106-113, 2001.

E. Manini and R. Danovaro, Synoptic determination of living/dead and active/dormant bacterial fractions in marine sediments, FEMS Microbiology Ecology, vol.55, issue.3, pp.416-423, 2006.
DOI : 10.1111/j.1574-6941.2005.00042.x

G. Mariotti and S. Fagherazzi, Modeling the effect of tides and waves on benthic biofilms, Journal of Geophysical Research: Biogeosciences, vol.29, issue.1, p.670, 2012.
DOI : 10.1016/S0378-3839(96)00006-3

D. S. Mclusky, The estuarine ecosystem, 1989.

J. J. Middelburg, C. Baranguet, H. T. Boschker, P. M. Herman, T. Moens et al., The fate of intertidal microphytobenthos carbon: an in situ 13C-labeling study, 2000.

M. M. Moeseneder and G. J. Herndl, Influence of turbulence on bacterial production in the sea, Limnology and Oceanography, vol.40, issue.8, pp.1466-1473, 1995.
DOI : 10.4319/lo.1995.40.8.1466

F. Orvain, P. G. Sauriau, C. Bacher, and M. Prineau, The influence of sediment cohesiveness on bioturbation effects due to Hydrobia ulvae on the initial erosion of intertidal sediments: A study combining flume and model approaches, Journal of Sea Research, vol.55, issue.1, pp.54-73, 2006.
DOI : 10.1016/j.seares.2005.10.002

URL : https://hal.archives-ouvertes.fr/hal-00148892

M. G. Pachiadaki, A. Kallionaki, A. Dählmann, G. J. De-lange, and K. A. Kormas, Diversity and Spatial Distribution of Prokaryotic Communities Along A Sediment Vertical Profile of A Deep-Sea Mud Volcano, Microbial Ecology, vol.143, issue.3???4, pp.655-668, 2011.
DOI : 10.1007/s00248-011-9855-2

R. J. Parkes, B. A. Cragg, and P. Wellsbury, Recent studies on bacterial populations and processes in subseafloor sediments: A review, Hydrogeology Journal, vol.8, issue.1, pp.11-28, 2000.
DOI : 10.1007/PL00010971

P. Pascal, C. Dupuy, P. Richard, C. Mallet, E. Armynot-du-châtelet et al., Seasonal variation in consumption of benthic bacteria by meio- and macrofauna in an intertidal mudflat, Limnology and Oceanography, vol.54, issue.4, pp.1048-1059, 2009.
DOI : 10.4319/lo.2009.54.4.1048

URL : https://hal.archives-ouvertes.fr/hal-00525685

D. M. Paterson and K. S. Black, Water Flow, Sediment Dynamics and Benthic Biology, p.695, 1999.
DOI : 10.1016/S0065-2504(08)60193-2

P. N. Polymenakou, S. Bertilsson, A. Tselepides, and E. G. Stephanou, Bacterial Community Composition in Different Sediments from the Eastern Mediterranean Sea: a Comparison of Four 16S Ribosomal DNA Clone Libraries, Microbial Ecology, vol.68, issue.3, pp.447-462, 2005.
DOI : 10.1007/s00248-005-0005-6

E. T. Porter, R. P. Mason, and L. P. Sanford, Effect of tidal resuspension on benthic-pelagic 701, 2010.

A. Pusceddu, C. Fiordelmondo, and R. Danovaro, Sediment Resuspension Effects on the Benthic Microbial Loop in Experimental Microcosms, Microbial Ecology, vol.61, issue.286, pp.602-613, 2005.
DOI : 10.1007/s00248-005-5051-6

K. Ravenschlag, K. Sahm, and R. Amann, Quantitative molecular analysis of the microbial 705 community in marine Arctic sediments (Svalbard) Applied and Environmental 706 Microbiology, pp.387-395, 2001.

F. Riemann and E. Helmke, Symbiotic Relations of Sediment-Agglutinating Nematodes and Bacteria in Detrital Habitats: The Enzyme-Sharing Concept, Marine Ecology, vol.119, issue.2, pp.93-113, 2002.
DOI : 10.1038/359139a0

L. Riemann and A. Winding, Community dynamics of free-living and particle-associated 710 bacterial assemblages during a freshwater phytoplankton bloom, Microbial Ecology, vol.42, pp.711-274, 2001.

W. Ritzrau and G. Graf, Increase of microbial biomass in the benthic turbidity zone of Kiel 713, 1992.

W. Ritzrau, Microbial activity in the benthic boundary layer: Small-scale distribution 715 and its relationship to the hydrodynamic regime, Journal of Sea Research, vol.36, pp.171-180, 1996.

S. S. Shapiro and M. B. Wilk, An analysis of variance test for normality (complete samples), Biometrika, vol.52, issue.3-4, pp.591-611, 1965.
DOI : 10.1093/biomet/52.3-4.591

J. L. Schmidt, J. W. Deming, P. A. Jumars, and R. G. Keil, Constancy of bacterial abundance in surficial marine sediments, Limnology and Oceanography, vol.43, issue.5, pp.976-982, 1998.
DOI : 10.4319/lo.1998.43.5.0976

J. Shimeta, C. L. Amos, S. E. Beaulieu, and S. L. Katz, Resuspension of benthic protists at 721 subtidal coastal sites with differing sediment composition, Marine Ecology Progress 722 Series, pp.103-115, 2003.

C. Ståhlberg, D. Bastviken, B. H. Svensson, and L. Rahm, Mineralisation of organic 724 matter in coastal sediments at different frequency and duration of resuspension, Estuarine, 725 Coastal and Shelf Science, pp.317-325, 2006.

E. Takahashi, J. Ledauphin, D. Goux, and F. Orvain, Optimising extraction of 727 extracellular polymeric substances (EPS) from benthic diatoms: comparison of the 728 efficiency of six EPS extraction methods. Marine and Freshwater Research, pp.1201-1210, 2009.

J. M. Teal, Energy Flow in the Salt Marsh Ecosystem of Georgia, Ecology, vol.43, issue.4, pp.614-624, 1962.
DOI : 10.2307/1933451

F. Thingstad, Control of bacterial growth in idealized food webs, In: Kirchman DL, p.731, 2000.

G. J. Underwood and D. M. Paterson, The importance of extracellular carbohydrate productionby marine epipelic diatoms, Advances in Botanical Research, vol.40, pp.183-240, 2003.
DOI : 10.1016/S0065-2296(05)40005-1

G. J. Underwood and J. Kromkamp, Primary Production by Phytoplankton and Microphytobenthos in Estuaries, Advances in Ecological Research, vol.29, pp.93-153, 1999.
DOI : 10.1016/S0065-2504(08)60192-0

F. C. Van-duyl, B. De-winder, A. J. Kop, and U. Wollenzien, Tidal coupling between 737 carbohydrate concentrations and bacterial activities in diatom-inhabited intertidal mudflats, p.738, 1999.

S. C. Wainright, Stimulation of Heterotrophic Microplankton Production by Resuspended Marine Sediments, Science, vol.238, issue.4834, pp.1710-1712, 1987.
DOI : 10.1126/science.238.4834.1710

S. C. Wainright, Sediment-to-water fluxes of particulate material and microbes by 742 resuspension and their contribution to the planktonic food web, Marine Ecology Progress 743 Series, pp.271-281, 1990.

J. H. Ward, Hierarchical Grouping to Optimize an Objective Function, Journal of the American Statistical Association, vol.58, issue.301, p.745, 1963.
DOI : 10.1007/BF02289263

E. J. Weerman, P. M. Herman, and J. Van-de-koppel, Top-down control inhibits spatial self-organization of a patterned landscape, Ecology, vol.40, issue.2, pp.487-495, 2011.
DOI : 10.1016/S0272-7714(02)00359-1

R. Wilms, H. Sass, B. Köpke, J. Köster, H. Cypionka et al., Specific bacterial3)ns 7.6 (1.5)ns 8.5 (0.1)a 127,8 (0.1)a 51,6 (0.2)a 21, Applied and Environmental Microbiology, vol.72413, issue.490, pp.2756-27647, 2006.