D. Abrol, Ecology, behaviour and management of social wasp, Vespa velutina smith (Hymenoptera: Vespidae ), attacking honeybee colonies, Korean J Apic, vol.9, pp.5-10, 1994.

M. Arca, A. Papachristoforou, F. Mougel, A. Rortais, K. Monceau et al., Defensive behaviour of Apis mellifera against Vespa velutina in France: Testing whether European honeybees can develop an effective collective defence against a new predator, Behavioural Processes, vol.106, pp.122-129, 2014.
DOI : 10.1016/j.beproc.2014.05.002

M. Archer, Taxonomy, distribution and nesting biology of species of the genera Provespa Ashmead and Vespa Linneaus (Hymenoptera, Vespidae), Entomol Mon Mag, vol.144, pp.69-101

M. Archer, Taxonomy, distribution and nesting biology of the species of the genus Paravespula or the Vespula vulgaris species group (Hym., Vespidae), Entomol Mon Mag, vol.144, p.5

M. Archer, The queen colony phase of vespine wasps (Hymenoptera, Vespidae), Insectes Sociaux, vol.57, issue.2, pp.133-145, 2010.
DOI : 10.1007/s00040-009-0063-8

M. Archer, Vespine wasps of the world. Behaviour, ecology and taxonomy of the vespinae, monograph series

M. Barbet-massin, Q. Rome, F. Muller, A. Perrard, C. Villemant et al., Climate change increases the risk of invasion by the Yellow-legged hornet, Biological Conservation, vol.157, pp.4-10, 2013.
DOI : 10.1016/j.biocon.2012.09.015

J. Beggs, E. Brockerhoff, J. Corley, M. Kenis, M. Masciocchi et al., Ecological effects and management of invasive alien Vespidae, BioControl, vol.1, issue.2, pp.505-526, 2011.
DOI : 10.1007/s10526-011-9389-z

B. Bradley, D. Blumenthal, R. Early, E. Grosholz, J. Lawler et al., Global change, global trade, and the next wave of plant invasions, Frontiers in Ecology and the Environment, vol.98, issue.1, pp.20-28, 2011.
DOI : 10.1890/110145.s1

D. Bunn, Notes on the nesting cycle of Dolichovespula sylvestris Scop. (Hym., Vespidae), Entomol Mon Mag, vol.118, pp.213-218, 1982.

M. Choi, S. Martin, and J. Lee, Distribution, spread, and impact of the invasive hornet Vespa velutina in South Korea, Journal of Asia-Pacific Entomology, vol.15, issue.3, pp.473-477, 2012.
DOI : 10.1016/j.aspen.2011.11.004

M. Choi, S. Lee, H. Suk, and J. Lee, , in South Korea, South Korea, pp.208-214, 2013.
DOI : 10.1111/1748-5967.12027

D. Dazhi and W. Yunzhen, A preliminary study on the biology of wasps Vespa velutina auraria Smith and Vespa tropica ducalis Smith (Hymenoptera: Vespidae), Zool Res, vol.10, pp.155-162, 1989.

S. Demichelis, A. Manimo, and M. Porporato, Trovato il primo nido di Vespavelutina a Vallecrosia (IM) (Communicato Stampa), 2013.

B. Donovan, Nest initiation by German and common wasp queens (Hymenoptera: Vespidae) and nest fate at Christchurch, New Zealand, New Zealand Journal of Zoology, vol.33, issue.2, pp.95-99, 1991.
DOI : 10.1080/03014223.1989.10422899

R. Edwards, Social wasps: their biology and control. Rentokil, East Grinstead, 1980.

K. Foster, J. Gulliver, and F. Ratnieks, Worker policing in the European hornet Vespa crabro, Insectes Sociaux, vol.49, issue.1, pp.41-44, 2002.
DOI : 10.1007/s00040-002-8277-z

A. Hastings, K. Cuddington, K. Davies, C. Dugaw, S. Elmendorf et al., The spatial spread of invasions: new developments in theory and evidence, Ecology Letters, vol.16, issue.1, pp.91-101, 2005.
DOI : 10.1111/j.1461-0248.2004.00687.x

J. Haxaire, J. Bouguet, and J. Tamisier, Vespa velutina Lepeletier, 1836, une redoutable nouveaut e pour la faune de France (Hym., Vespidae), 0194.

I. Ib-a~-nez, J. Diez, L. Miller, J. Olden, C. Sorte et al., Integrated assessment of biological invasions, Ecological Applications, vol.24, issue.1, pp.25-37, 2014.
DOI : 10.1126/science.277.5325.494

M. Inoue, J. Yokoyama, and K. Tsuchida, Colony growth and reproductive ability of feral nests of the introduced bumblebee Bombus terrestris in northern Japan, Insectes Sociaux, vol.57, issue.1, pp.29-38, 2010.
DOI : 10.1007/s00040-009-0047-8

C. Janet, Observations sur les gu^ epes, C. Naud, 1903.

C. Jung, M. Kang, D. Kim, and H. Lee, Vespid wasps (Hymenoptera: Vespidae) occurring around apiaries in Andong, Korea. I. taxonomy and life history, Korean J Apic, vol.22, pp.53-62, 2007.

C. Jung, D. Kim, H. Lee, and H. Baek, Some biological characteristics of a new honeybee pest, Vespa velutina nigrithorax Buysson 1905 (Hymenoptera: Vespidae), Korean J Apic, vol.24, pp.61-65, 2008.

T. Kadoya, H. Ishii, R. Kikuchi, S. Suda, and I. Washitani, Using monitoring data gathered by volunteers to predict the potential distribution of the invasive alien bumblebee Bombus terrestris, Biological Conservation, vol.142, issue.5, pp.1011-1017, 2009.
DOI : 10.1016/j.biocon.2009.01.012

J. Kim, M. Choi, and T. Moon, Occurrence of Vespa velutina Lepeletier from Korea, and a revised key for Korean Vespa species (Hymenoptera: Vespidae), Entomological Research, vol.34, issue.2, p.112, 2006.
DOI : 10.1111/j.1748-5967.2006.00018.x

M. Kuo and W. Yeh, Ecological studies on Vespa basalis Smith, Vespa velutina flavitarsus Sonan and Tropica pseudosoror Vecht. (Study on Vespidae in Taiwan II), J Natl Chiayi Inst Agric, vol.11, pp.95-106, 1985.

O. Latter, A reason for the order in which the queen wasp constructs the cells of the comb, a method for computing the number of cells in a ctHnb, and an instance of the limitation of the instincts of wasps, Proc R Entomol Soc Lond, vol.10, pp.74-78, 1935.

S. Martin, Weight changes in adult hornets,Vespa affinis (Hymenoptera: Vespidae), Insectes Sociaux, vol.34, issue.4, pp.363-368, 1993.
DOI : 10.1007/BF01253899

M. Matsuura, Nesting habit of several species of the genus Vespa in Formosona, Kontyu, vol.41, pp.286-293, 1973.

M. Matsuura, Comparative biology of the five Japanese species of the genus Vespa (Hymenoptera, Vespidae ), pp.1-131, 1984.

M. Matsuura and S. Yamane, Biology of the Vespine Wasps, 1990.
DOI : 10.1007/978-3-642-75230-8

K. Monceau, O. Bonnard, and T. , Chasing the queens of the alien predator of honeybees: A water drop in the invasiveness ocean, Open Journal of Ecology, vol.02, issue.04, pp.183-191, 2012.
DOI : 10.4236/oje.2012.24022

K. Monceau, M. Arca, L. Lepr^-etre, F. Mougel, O. Bonnard et al., Native Prey and Invasive Predator Patterns of Foraging Activity: The Case of the Yellow-Legged Hornet Predation at European Honeybee Hives, PLoS ONE, vol.84, issue.6, p.66492, 2013.
DOI : 10.1371/journal.pone.0066492.t003

M. Nakamura and S. Sonthichai, Nesting habits of some hornet species (Hymenoptera, Vespidae) in Northern Thailand, Kasetsart J Nat Sci, vol.38, 0196.

D. Paini and J. Roberts, Commercial honey bees () reduce the fecundity of an Australian native bee (), Biological Conservation, vol.123, issue.1, pp.103-112, 2005.
DOI : 10.1016/j.biocon.2004.11.001

A. Perrard, J. Haxaire, A. Rortais, and C. Villemant, Observations on the colony activity of the Asian Hornet Vespa velutina Lepeletier 1836 (hymenoptera: Vespidae: Vespinae ) in France, Ann Soc Entomol Fr, pp.45-55, 2009.

A. Perrard, C. Villemant, J. Carpenter, and M. Baylac, Differences in caste dimorphism among three hornet species (Hymenoptera: Vespidae): forewing size, shape and allometry, Journal of Evolutionary Biology, vol.42, issue.7, pp.1389-1398, 2012.
DOI : 10.1111/j.1420-9101.2012.02527.x

R. Team, R: a language and environment for statistical computing. R Foundation for Statistical Computing, 2014.

N. Ranabhat and A. Tamrakar, Study on seasonal activity of predatory wasps attacking honeybee Apis cerana Fab. colonies in Southern belt of Kaski district, Nepal, J Nat Hist Mus, vol.23, pp.125-128, 2009.

J. Rasplus, C. Villemant, M. Paiva, G. Delvare, and A. Roques, In: Arthropod Invasions in Europe, Hymenoptera. Chapter, vol.12, pp.669-776, 2010.

Q. Rome, F. Muller, O. Gargominy, and C. Villemant, Bilan 2008 De l'invasion de Vespa velutina Lepeletier en France (Hymenoptera: Vespidae), Bull Soc Entomol Fr, vol.114, pp.297-302, 2009.

Q. Rome, A. Perrard, F. Muller, and C. Villemant, Monitoring and control modalities of a honeybee predator, the yellow-legged hornet Vespa velutina nigrithorax (Hymenoptera: Vespidae). Aliens, pp.31-38, 2011.

A. Roques, W. Rabitsch, J. Rasplus, C. Lopez-vaamonde, W. Nentwig et al., Alien Terrestrial Invertebrates of Europe, Handbook of Alien Species in Europe , Invading Nature ? Springer Series in Invasion Ecology, pp.63-79, 2009.
DOI : 10.1007/978-1-4020-8280-1_5

A. Rortais, C. Villemant, O. Gargominy, Q. Rome, J. Haxaire et al., A new enemy of honeybees in Europe: the Asian Hornet Vespa velutina. In: Ed. by Settele J. Atlas of biodiversity risks ? from Europe to the Globe, from stories to maps, p.181, 2010.

L. Sheng and L. Xing, Species and identification of the sting Wasps distributed in the City Zone of Hangzhou, J Huzhou Vocat Tech Coll, vol.3, pp.85-87, 2005.

J. Sonan, On Vespa from formosa, Trans Nat Hist Soc Formosa, vol.19, pp.136-149, 1929.

J. Spradbery, Wasps: an account of the biology and natural history of solitary and social wasps with particular reference to those of the British Isles, 1973.

K. Tan, S. Radloff, J. Li, H. Hepburn, M. Yang et al., Bee-hawking by the wasp, Vespa velutina, on the honeybees Apis cerana and A. mellifera, Naturwissenschaften, vol.14, issue.6, pp.469-472, 2007.
DOI : 10.1007/s00114-006-0210-2

C. Thomas, The European Wasp (Vespula germanica Fab.) in New Zealand, NZ Dep Sci Ind Res Inf Ser, vol.27, pp.1-74, 1960.

J. Van-der-vecht, The Vespinae of the Indo-Malayan and Papuan areas (Hymenoptera: Vespidae), 1957.

C. Villemant, J. Haxaire, and J. Streito, Premier bilan de l'invasion de Vespa velutina lepeletier en France (Hymenoptera , Vespidae), Bull Soc Entomol Fr, vol.111, p.535, 2006.
URL : https://hal.archives-ouvertes.fr/hal-00172890

C. Villemant, M. Barbet-massin, A. Perrard, F. Muller, O. Gargominy et al., Predicting the invasion risk by the alien bee-hawking Yellow-legged hornet Vespa velutina nigrithorax across Europe and other continents with niche models, Biological Conservation, vol.144, issue.9, pp.2142-2150, 2011.
DOI : 10.1016/j.biocon.2011.04.009

C. Villemant, F. Muller, Q. Rome, A. Perrard, M. Barbet-massin et al., Estimating the potential range expansion and environmental impact of the invasive beehawking hornet, Vespa velutina nigrithorax. In: In silico bees, pp.269-287, 2014.

T. Wenseleers, H. Helanter?-a, A. Hart, and F. Ratnieks, Worker reproduction and policing in insect societies: an ESS analysis, Journal of Evolutionary Biology, vol.29, issue.5, pp.1035-1047, 2004.
DOI : 10.1111/j.1420-9101.2004.00751.x

T. Wenseleers, N. Badcock, K. Erven, A. Tofilski, F. Nascimento et al., A TEST OF WORKER POLICING THEORY IN AN ADVANCED EUSOCIAL WASP, VESPULA RUFA, Evolution, vol.128, issue.6, pp.59-1306, 2005.
DOI : 10.1007/BF00343205

S. Yamane, On the collection technique of vespine nests, based chiefly on practices through a survey in Taiwan from 1972 to 1974 (Hymenoptera: Vespidae), Seibutsu Kyozai, vol.12, pp.42-59, 1977.

S. Yamane, A huge nest of Vespa basalis in Taiwan (Hymenoptera: Vespidae), Chin J Entomol, vol.12, pp.1-11, 1992.