M. A. Allison, M. T. Lee, A. S. Ogston, and R. C. Aller, Origin of Amazon mudbanks along the northeastern coast of South America, Marine Geology, vol.163, issue.1-4, pp.241-256, 2000.
DOI : 10.1016/S0025-3227(99)00120-6

D. M. Alongi, Intertidal zonation and seasonality of meiobenthos in tropical mangrove estuaries, Marine Biology, vol.42, issue.3, pp.447-458, 1987.
DOI : 10.1007/BF00409574

D. M. Alongi, The role of soft-bottom benthic communities in tropical mangrove and coral reef ecosystems, Revue Aquatic Sciences, vol.1, pp.243-280, 1989.

E. J. Anthony, A. Gardel, N. Gratiot, C. Proisy, M. A. Allison et al., The Amazon-influenced muddy coast of South America: A review of mud-bank???shoreline interactions, Earth-Science Reviews, vol.103, issue.3-4, pp.99-121, 2010.
DOI : 10.1016/j.earscirev.2010.09.008

URL : https://hal.archives-ouvertes.fr/insu-00562710

M. Balsamo, G. Albertelli, V. U. Ceccherelli, R. Coccioni, M. A. Colangelo et al., Meiofauna of the Adriatic Sea: present knowledge and future perspectives, Meiofauna of the Adriatic Sea: current state of knowledge and future perspectives, pp.45-63, 2010.
DOI : 10.1016/j.energy.2004.03.051

M. Balsamo, F. Semprucci, F. Frontalini, and R. Coccioni, Meiofauna as a Tool for Marine Ecosystem Biomonitoring, pp.77-104, 2012.
DOI : 10.5772/34423

S. J. Blott and K. Pye, GRADISTAT: a grain size distribution and statistics package for the analysis of unconsolidated sediments, Earth Surface Processes and Landforms, vol.58, issue.11, pp.1237-1248, 2001.
DOI : 10.1002/esp.261

J. Bohórquez, S. Papaspyrou, M. Yúfera, S. A. Van-bergeijk, E. García-robledo et al., Effects of green macroalgal blooms on the meiofauna community structure in the Bay of C??diz, Marine Pollution Bulletin, vol.70, issue.1-2, pp.10-17, 2013.
DOI : 10.1016/j.marpolbul.2013.02.002

G. Boucher and P. J. Lambshead, Ecological Biodiversity of Marine Nematodes in Samples from Temperate, Tropical, and Deep-Sea Regions, Conservation Biology, vol.9, issue.6, pp.1594-1604, 1995.
DOI : 10.1046/j.1523-1739.1995.09061594.x

G. Chinnadurai and O. J. Fernando, Meiofauna of mangroves of the southeast coast of India with special reference to the free-living marine nematode assemblage, Estuarine, Coastal and Shelf Science, vol.72, issue.1-2, pp.329-336, 2007.
DOI : 10.1016/j.ecss.2006.11.004

B. C. Coull, Role of meiofauna in estuarine soft-bottom habitats*, Austral Ecology, vol.228, issue.Suppl., pp.327-343, 1999.
DOI : 10.1046/j.1442-9993.1999.00979.x

J. P. Debenay, D. Guiral, and M. Parra, Ecological Factors Acting on the Microfauna in Mangrove Swamps. The Case of Foraminiferal Assemblages in French Guiana, Estuarine, Coastal and Shelf Science, vol.55, issue.4, pp.509-533, 2002.
DOI : 10.1006/ecss.2001.0906

S. Dittmann, Zonation of benthic communities in a tropical tidal flat of north-east Australia, Journal of Sea Research, vol.43, issue.1, pp.33-51, 2000.
DOI : 10.1016/S1385-1101(00)00004-6

G. Y. Du, M. Son, M. Yun, S. An, and I. K. Chung, Microphytobenthic biomass and species composition in intertidal flats of the Nakdong River estuary, Korea, Estuarine, Coastal and Shelf Science, vol.82, issue.4, pp.663-672, 2009.
DOI : 10.1016/j.ecss.2009.03.004

C. Dupuy, C. Mallet, K. Guizien, H. Montanié, M. Bréret et al., Sequential resuspension of biofilm components (viruses, prokaryotes and protists) as measured by erodimetry experiments in the Brouage mudflat (French Atlantic coast), Journal of Sea Research, vol.92, pp.56-65, 2014.
DOI : 10.1016/j.seares.2013.12.002

URL : https://hal.archives-ouvertes.fr/hal-01104286

C. Heip, M. Vincx, and G. Vranken, The ecology of marine nematodes, Oceanography and Marine Biology Annual Review, vol.23, pp.399-489, 1985.

C. Heip, R. Huys, and R. Alkemade, Community structure and functional roles of meiofauna in the North Sea, Netherlands Journal of Aquatic Ecology, vol.72, issue.6, pp.31-41, 1992.
DOI : 10.1007/BF02298026

O. Herlory, J. M. Guarini, P. Richard, and G. F. Blanchard, Microstructure of microphytobenthic biofilm and its spatio-temporal dynamics in an intertidal mudflat (Aiguillon Bay, France), Marine Ecology Progress Series, vol.282, pp.33-44, 2004.
DOI : 10.3354/meps282033

URL : https://hal.archives-ouvertes.fr/hal-00093955

S. C. Kapusta, N. L. Würdig, C. E. Bemvenuti, and C. P. Ozorio, Meiofauna structure in Tramandaí-Armazém estuary (South of Brazil), Acta Limnologica Brasiliensia, vol.17, pp.349-359, 2005.

?. Lalana, R. Rueda, and F. Gosselck, Investigations of the Benthos of Mangrove Coastal Lagoons in Southern Cuba, Internationale Revue der gesamten Hydrobiologie und Hydrographie, vol.33, issue.6, pp.779-794, 1986.
DOI : 10.1002/iroh.19860710605

C. Lavergne, L. Beaugeard, C. Dupuy, C. Courties, and H. Agogué, An efficient and rapid method for the enumeration of heterotrophic prokaryotes in coastal sediments by flow cytometry, Journal of Microbiological Methods, vol.105, pp.31-38, 2014.
DOI : 10.1016/j.mimet.2014.07.002

C. J. Lorenzen, A method for the continuous measurement of in vivo chlorophyll concentration. Deep-Sea Research and Oceanography Abstract 13, pp.223-227, 1966.

D. Marie, F. Partensky, D. Vaulot, and C. Brussaard, Enumeration of Phytoplankton, Bacteria, and Viruses in Marine Samples, 2001.
DOI : 10.1002/0471142956.cy1111s10

J. M. Martinez, J. L. Guyot, N. Filizola, and F. Sondag, Increase in suspended sediment discharge of the Amazon River assessed by monitoring network and satellite data, CATENA, vol.79, issue.3, pp.257-264, 2009.
DOI : 10.1016/j.catena.2009.05.011

URL : https://hal.archives-ouvertes.fr/ird-00394090

J. J. Middelburg, C. Barranguet, H. T. Boschker, P. M. Herman, T. Moens et al., C-labeling study, Limnology and Oceanography, vol.45, issue.6, pp.1224-1234, 2000.
DOI : 10.4319/lo.2000.45.6.1224

T. Moens and M. Vincx, Observations on the Feeding Ecology of Estuarine Nematodes, Journal of the Marine Biological Association of the United Kingdom, vol.28, issue.01, pp.211-227, 1997.
DOI : 10.2307/1540955

T. Moens, C. Luyten, J. J. Middelburg, P. M. Herman, and M. Vincx, Tracing organic matter sources of estuarine tidal flat nematodes with stable carbon isotopes, Marine Ecology Progress Series, vol.234, pp.127-137, 2002.
DOI : 10.3354/meps234127

V. O. Mokievsky, A. V. Tchesunovb, A. A. Udalova, N. Duy, and T. , Quantitative distribution of meiobenthos and the structure of the free-living nematode community of the mangrove intertidal zone in Nha Trang bay (Vietnam) in the South China Sea, Russian Journal of Marine Biology, vol.37, issue.4, 2011.
DOI : 10.1134/S1063074011040109

P. A. Montagna, G. F. Blanchard, and A. Dinet, Effect of production and biomass of intertidal microphytobenthos on meiofaunal grazing rates, Journal of Experimental Marine Biology and Ecology, vol.185, issue.2, pp.149-165, 1995.
DOI : 10.1016/0022-0981(94)00138-4

F. Orvain, K. Guizien, S. Lefebvre, M. Bréret, and C. Dupuy, Relevance of macrozoobenthic grazers to understand the dynamic behaviour of sediment erodibility and microphytobenthos resuspension in sunny summer conditions, Journal of Sea Research, vol.92, pp.46-55, 2014.
DOI : 10.1016/j.seares.2014.03.004

P. Y. Pascal, C. Dupuy, P. Richard, A. G. Haubois, and N. Niquil, Influence of environment factors on bacterial ingestion rate of the deposit-feeder Hydrobia ulvae and comparison with meiofauna, Journal of Sea Research, vol.60, issue.3, pp.151-156, 2008.
DOI : 10.1016/j.seares.2008.05.003

URL : https://hal.archives-ouvertes.fr/hal-01248039

P. Y. Pascal, C. Dupuy, P. Richard, J. Rzeznik-orignac, and N. Niquil, Bacterivory of a mudflat nematode community under different environmental conditions, Marine Biology, vol.5, issue.4, pp.671-682, 2008.
DOI : 10.1007/s00227-008-0960-9

URL : https://hal.archives-ouvertes.fr/hal-00293542

P. Y. Pascal, C. Dupuy, P. Richard, C. Mallet, E. Armynot-du-chatelet et al., Seasonal variation in consumption of benthic bacteria by meio- and macrofauna in an intertidal mudflat, Limnology and Oceanography, vol.54, issue.4, pp.1048-1059, 2009.
DOI : 10.4319/lo.2009.54.4.1048

URL : https://hal.archives-ouvertes.fr/hal-00525685

R. G. Perkins, C. Honeywill, M. Consalvey, H. A. Austin, T. J. Tolhurst et al., Changes in microphytobenthic chlorophyll a and EPS resulting from sediment compaction due to de-watering: opposing patterns in concentration and content, Continental Shelf Research, vol.23, issue.6, pp.575-586, 2003.
DOI : 10.1016/S0278-4343(03)00006-2

J. L. Pinckney, K. R. Carman, S. E. Lumsden, and S. N. Hymel, Microalgal-meiofaunal trophic relationships in muddy intertidal estuarine sediments, Aquatic Microbial Ecology, vol.31, pp.99-108, 2003.
DOI : 10.3354/ame031099

H. M. Platt and R. M. Warwick, The significance of free living nematodes to the littoral ecosystem The shore environment: 2.Ecosystems, Systematics association special, pp.729-759, 1980.

J. Rzeznik-orignac, D. Fichet, and G. Boucher, Spatio-temporal structure of the nematode assemblages of the Brouage mudflat (Marennes Ol??ron, France), Estuarine, Coastal and Shelf Science, vol.58, issue.1, pp.77-88, 2003.
DOI : 10.1016/S0272-7714(03)00061-1

J. Rzeznik-orignac and D. Fichet, Experimental estimation of assimilation rates of meiofauna feeding on C-14-labelled benthic diatoms, Journal of Experimental Marine Biology and Ecology, vol.432, pp.179-185, 2012.

F. Semprucci, P. Colantoni, G. Baldelli, M. Rocchi, and M. Balsamo, The distribution of meiofauna on back-reef sandy platforms in the Maldives (Indian Ocean) Marine Ecology: An evolutionary perspective, pp.592-607, 2010.

F. Semprucci, P. Colantoni, C. Sbrocca, G. Baldelli, M. Rocchi et al., Meiofauna in sandy back-reef platforms differently exposed to the monsoons in the Maldives (Indian Ocean), Journal of Marine Systems, vol.87, issue.3-4, pp.208-215, 2011.
DOI : 10.1016/j.jmarsys.2011.04.002

J. L. Schmidt, J. W. Deming, P. A. Jumars, and R. G. Keil, Constancy of bacterial abundance in surficial marine sediments, Limnology and Oceanography, vol.43, issue.5, pp.976-982, 1998.
DOI : 10.4319/lo.1998.43.5.0976

P. Schwinghamer, Characteristic Size Distributions of Integral Benthic Communities, Canadian Journal of Fisheries and Aquatic Sciences, vol.38, issue.10, pp.1255-1263, 1981.
DOI : 10.1139/f81-167

K. Soetaert, M. Vincx, J. Wittoeck, and M. Tulkens, Meiobenthic distribution and nematode community structure in five European estuaries, Hydrobiologia, vol.18, issue.283, pp.185-206, 1995.
DOI : 10.1007/BF00008580

T. J. Tolhurst, E. C. Defew, and A. Dye, Lack of correlation between surface macrofauna, meiofauna, erosion threshold and biogeochemical properties of sediments within an intertidal mudflat and mangrove forest, Hydrobiologia, vol.189, issue.3, pp.1-13, 2010.
DOI : 10.1007/s10750-010-0311-y

G. J. Underwood and J. Kromkamp, Primary Production by Phytoplankton and Microphytobenthos in Estuaries, Advances in Ecological Research, vol.29, pp.93-153, 1999.
DOI : 10.1016/S0065-2504(08)60192-0

G. J. Underwood and D. M. Paterson, The importance of extracellular carbohydrate productionby marine epipelic diatoms, Advances in Botanical Research, vol.40, pp.183-240, 2003.
DOI : 10.1016/S0065-2296(05)40005-1

G. J. Underwood, Microphytobenthos and phytoplankton in the Severn estuary, UK: Present situation and possible consequences of a tidal energy barrage, Marine Pollution Bulletin, vol.61, issue.1-3, pp.83-91, 2010.
DOI : 10.1016/j.marpolbul.2009.12.015

F. C. Van-duyl, B. De-winder, A. J. Kop, and U. Wollenzien, Tidal coupling between carbohydrate concentrations and bacterial activities in diatom-inhabited intertidal mudflats, Marine Ecology Progress Series, vol.191, pp.19-32, 1999.
DOI : 10.3354/meps191019

S. Vanhove, M. Vincx, D. V. Gansbeke, W. Gijselinck, and D. Schram, The meiobenthos of five mangrove vegetation types in Gazi Bay, Kenya, Hydrobiologia, vol.41, issue.1-3, pp.99-108, 1992.
DOI : 10.1007/BF00008208

D. Van-oevelen, K. Soetaert, J. J. Middelburg, P. M. Herman, L. Moodley et al., Carbon flows through a benthic food web: Integrating biomass, isotope and tracer data, Journal of Marine Research, vol.64, issue.3, pp.453-482, 2006.
DOI : 10.1357/002224006778189581

R. M. Warwick and J. H. Price, Ecological and metabolic studies on free-living nematodes from an estuarine mud-flat, Estuarine and Coastal Marine Science, vol.9, issue.3, pp.257-271, 1979.
DOI : 10.1016/0302-3524(79)90039-2

W. Wieser, Die Beziehungen zwischen Mundho¨hlengestaltMundho¨hlengestalt, Erna¨hrungsweiseErna¨hrungsweise und Vorkommen bei freilebenden marinen Nematoden. Eine o¨kologischo¨kologisch-morphologische Studie, Arkiv fur Zoologi, vol.4, pp.439-484, 1953.

W. Wieser, BENTHIC STUDIES IN BUZZARDS BAY II. THE MEIOFAUNA1, Limnology and Oceanography, vol.5, issue.2, pp.121-137, 1960.
DOI : 10.4319/lo.1960.5.2.0121

R. Wollast, Physico-chemical models of marine sediment Proceedings of the international seminar on the environmental aspects of dredging activities, pp.191-204, 1989.