D. Aedo, Esporas de algas rojas coalescentes y no coalescentes: diferencias 472 funcionales y factores ambientales afectado la coalescencia, p.473, 2007.

N. L. Andrew and R. M. Viejo, Effects of wave exposure and intraspecific density on the 476 growth and survivorship of Sargassum muticum (Sargassaceae: Phaeophyta), 1998.

M. Biedrzycki, T. Jilany, S. Dudley, and H. Bais, Root exudates mediate kin recognition 483 in plants, Communicative & Integrated Biology, vol.3, pp.1-8, 2010.

B. M. Bolker, M. E. Brooks, C. J. Clarck, S. W. Geange, J. R. Poulsen et al., Generalized linear mixed models: a practical guide for ecology and evolution. 488 Trends in Ecology and Evolution, pp.127-135, 2008.

B. R. Broitman, S. A. Navarrete, F. Smith, and S. D. Gaines, Geographic variation of 490 southeastern Pacific intertidal communities, Marine Ecology Progress Series, vol.224, pp.491-512, 2001.

P. Camus, Recruitment of the intertidal kelp Lessonia nigrescens Bory in northern Chile: successional constraints and opportunities, Journal of Experimental Marine Biology and Ecology, vol.184, issue.2, pp.184-171, 1994.
DOI : 10.1016/0022-0981(94)90003-5

M. J. Crawley, Statistics: an introduction using R, 2005.
DOI : 10.1002/9781119941750

P. K. Dayton, Experimental Evaluation of Ecological Dominance in a Rocky Intertidal Algal Community, Ecological Monographs, vol.45, issue.2, pp.137-159, 1974.
DOI : 10.2307/1942404

P. K. Dayton, V. Currie, T. Gerrodette, B. Keller, R. Ronsenthal et al., Patch 499 dynamics and stability of some southern California kelp communities, pp.253-289, 1984.

H. De-kroon, Ecology: how do roots interact? Science, pp.1562-1563, 2007.

G. Gerlach, A. Hodgins-davis, B. Macdonald, and R. Hannah, Benefits of kin association: related and familiar zebrafish larvae (Danio rerio) show improved growth, Behavioral Ecology and Sociobiology, vol.3, issue.Suppl 1, pp.1765-1770, 2007.
DOI : 10.1007/s00265-007-0409-z

A. V. González and B. Santelices, Coalescence and chimerism in Codium (Chlorophyta) from central Chile, Phycologia, vol.47, issue.5, pp.468-476, 2008.
DOI : 10.2216/07-86.1

B. Santelices, Identification of cryptic species in the Lessonia nigrescens 512 complex (Phaeophyceae, Laminariales), Journal of Phycology, vol.48, pp.1153-1165, 2012.

A. S. Griffin and S. A. West, Kin selection: fact and fiction. Trends in Ecology and 514 Evolution, pp.15-21, 2002.
DOI : 10.1016/s0169-5347(01)02355-2

C. E. Grueber, S. Nakagawa, R. J. Laws, and I. G. Jamieson, Multimodel inference in 516 ecology and evolution: challenges and solutions, Journal of Evolutionary Biology, vol.24, pp.517-699, 2011.

W. D. Hamilton, The genetical evolution of social behaviour. I, Journal of Theoretical Biology, vol.7, issue.1, pp.1-16, 1964.
DOI : 10.1016/0022-5193(64)90038-4

M. S. Johnson and R. Black, Chaotic genetic patchiness in an intertidal limpet, p.523, 1982.

M. Lynch and K. Ritland, Estimation of pairwise relatedness with molecular markers, Genetics, vol.527, issue.152, pp.1753-1766, 1999.

T. Malm and . Kautskyl, Are Bladderwrack (Fucus vesiculosus L.) holdfasts that 529 support several fronds composed of one or several genetic individuals? Aquatic 530, Botany, vol.80, pp.221-226, 2004.

E. Martínez and B. Santelices, Selective mortality on haploid and diploid microscopic stages of Lessonia nigrescens Bory (Phaeophyta, Laminariales), Journal of Experimental Marine Biology and Ecology, vol.229, issue.2, pp.219-239, 1998.
DOI : 10.1016/S0022-0981(98)00063-X

E. Martínez, L. Cárdenas, and R. Pinto, Recovery and genetic diversity of the intertidal 535, 2003.

O. Nachtomy, A. Shavit, and J. Smith, Leibnizian organisms, nested individuals, and units of selection, Theory in Biosciences, vol.17, issue.2, pp.205-230, 2002.
DOI : 10.1007/s12064-002-0020-9

W. A. Nelson, Life history and growth in culture of the endemic New Zealand kelp 544, 2004.

J. Lessonia-variegata, Agardh in response to differing regimes of temperature, 545 photoperiod and light, Journal of Applied Phycology, vol.17, pp.23-28

A. Novoplansky, Picking battles wisely: plant behaviour under competition. Plant, 547 Cell and Environment, pp.726-741, 1998.

F. P. Ojeda and B. Santelices, Ecological dominance of Lessonia nigrescens 549 (Phaeophyta) in central Chile, Marine Ecology Progress Series, vol.16, pp.83-91, 1984.

R. Paine, BENTHIC MACROALGAL COMPETITION: COMPLICATIONS AND CONSEQUENCES, Journal of Phycology, vol.26, issue.1, pp.12-17, 1990.
DOI : 10.1111/j.0022-3646.1990.00012.x

R. Peakall and P. Smouse, genalex 6: genetic analysis in Excel. Population genetic software for teaching and research, Molecular Ecology Notes, vol.35, issue.1, pp.288-295, 2006.
DOI : 10.1046/j.1365-2540.1999.00518.x

J. W. Pepper and M. D. Herron, Does Biology Need an Organism Concept? Biological 555 Reviews, pp.621-627, 2008.

C. Team, R: a language and environment for statistical computing. R 557 Foundation for Statistical Computing, 2013.

K. Ritland, Estimators for pairwise relatedness and individual inbreeding coefficients, Genetical Research, vol.107, issue.02, pp.175-185, 1996.
DOI : 10.1038/hdy.1991.63

D. Rodriguez, O. M. Vásquez, and J. A. , Coalescence in wild organisms of the intertidal population of Lessonia berteroana in northern Chile: management and sustainability effects, Journal of Applied Phycology, vol.24, issue.2, pp.1115-1122, 2014.
DOI : 10.1007/s10811-013-0121-5

B. Santelices, Bases biológicas para el manejo de Lessonia nigrescens 564 (Phaeophyta Laminariales) en Chile Central, Monografías Biológicas, vol.2, pp.135-150, 1982.

B. Santelices, How many kind of individuals are there? Trends in Ecology and 566 Evolution, pp.152-155, 1999.

B. Santelices, Implications of clonal and chimeric-type thallus organization on 568 seaweed farming and harvesting, Journal of Applied Phycology, vol.13, issue.2, pp.153-160, 2001.
DOI : 10.1023/A:1011196327826

B. Santelices, A comparison of ecological responses among aclonal (unitary), clonal 570 and coalescing macroalgae, Journal of Experimental Marine Biology and Ecology, vol.571, pp.26-300, 2004.

B. Santelices and D. Aedo, Group Recruitment and Early Survival of Mazzaella Laminarioides, Journal of Applied Phycology, vol.143, issue.399, pp.583-589, 2006.
DOI : 10.1007/s10811-006-9067-1

B. Santelices and L. Alvarado, Demographic consequences of coalescence in sporeling 578 populations of Mazzaella laminaroides (Gigartinales, Rhodophyta) Journal of, p.579, 2008.

B. Santelices, J. Correa, I. Meneses, D. Aedo, and D. Varela, Sporeling coalescence and 581 intraclonal variation in Gracilaria chilensis, p.582, 1996.

B. Santelices and F. P. Ojeda, Recruitment, growth and survival of Lessonia nigrescens (Phaeophyta) at various tidal levels in exposed habitats of central Chile, Marine Ecology Progress Series, vol.19, pp.73-82, 1984.
DOI : 10.3354/meps019073

K. A. Selkoe, S. D. Gaines, J. E. Caselle, and R. R. Warner, CURRENT SHIFTS AND KIN AGGREGATION EXPLAIN GENETIC PATCHINESS IN FISH RECRUITS, Ecology, vol.87, issue.12, pp.3082-3094, 2006.
DOI : 10.1890/0012-9658(2006)87[3082:CSAKAE]2.0.CO;2

H. Steen and R. Scrosati, Intraspecific competition in Fucus serratus and F. evanescens 592 (Phaeophyceae: Fucales) germings: effects of settlement density, nutrient 593 concentration, and temperature, Marine Biology, vol.144, pp.61-70, 2004.

T. Torimaru, N. Tani, Y. Tsumura, N. Nishimura, and N. Tomaru, EFFECTS OF KIN-STRUCTURED SEED DISPERSAL ON THE GENETIC STRUCTURE OF THE CLONAL DIOECIOUS SHRUB ILEX LEUCOCLADA, Evolution, vol.28, issue.6, pp.1289-1300, 2007.
DOI : 10.2307/3236402

R. L. Vadas, A. W. Wright, and S. L. Miller, Recruitment of Ascophyllum nodosum: wave action as a source of mortality, Marine Ecology Progress Series, vol.61, pp.263-272, 1990.
DOI : 10.3354/meps061263

R. L. Vadas, S. Johnson, and T. A. Norton, Recruitment and mortality of early post-settlement stages of benthic algae, British Phycological Journal, vol.8, issue.3, pp.331-351, 1992.
DOI : 10.2307/1942623

C. Van-oosterhout, W. F. Hutchisnson, D. P. Willis, and P. Shipley, micro-checker: software for identifying and correcting genotyping errors in microsatellite data, Molecular Ecology Notes, vol.8, issue.3, pp.535-538, 2004.
DOI : 10.1046/j.1365-294x.1998.00477.x

J. A. Vásquez, Variables morfométricas y relaciones morfológicas de Lessonia 605 trabeculata Villouta & Santelices, 1984.

J. A. Vásquez, Lessonia trabeculata, a Subtidal Bottom Kelp in Northern Chile: A Case Study for a Structural and Geographical Comparison, Coastal communities of 609, 1992.
DOI : 10.1016/B978-0-08-092567-7.50011-8

J. A. Vásquez, Ecological Effects of Brown Seaweed Harvesting, Botanica Marina, vol.38, issue.1-6, pp.611-251, 1995.
DOI : 10.1515/botm.1995.38.1-6.251

J. A. Vásquez and A. Buschmann, Herbivory-kelp interactions in subtidal Chilean 613 communities: a review, Revista Chilena de Historia Natural, vol.70, pp.41-52, 1997.

J. A. Vásquez and B. Santelices, Comunidades de macroinvertebrados en discos de 615 adhesion de Lessonia nigrescens en Chile central, Revista Chilena de Historia 616 Natural, pp.131-154, 1984.

J. A. Vásquez and B. Santelices, Ecological effects of harvesting Lessonia (Laminariales, p.618, 1990.

J. A. Vásquez, F. Tala, J. M. Vega, S. Zuñiga, M. Edding et al., Bases 620 ecológicas y evaluación de usos alternativos para el manejo de praderas de algas 621 pardas de la III y IV regiones, 2008.

D. Veliz, P. Duchesne, E. Bourget, and L. Bernatchez, Genetic evidence for kin 623 aggregation in the intertidal acorn barnacle (Semibalanus balanoides) Molecular 624 Ecology, pp.4193-4202, 2006.

M. Venegas, F. Tala, E. Fonck, and J. A. Vásquez, Sporangial sori on stipes of Lessonia 626 nigrescens Bory (Laminareacea, Phaeophyta): a high frequency phenomenon in 627 intertidal populations of northern Chile, Botanica Marina, vol.35, pp.573-578, 1992.

T. Wernberg, G. A. Kendrick, and B. D. Toohey, Modification of the physical environment by an Ecklonia radiata (Laminariales) canopy and implications for associated foliose algae, Aquatic Ecology, vol.126, issue.205, pp.419-430, 2005.
DOI : 10.1007/s10452-005-9009-z

T. Wernberg, Holdfast aggregation in relation to morphology, age, attachment and drag for the kelp Ecklonia radiata, Aquatic Botany, vol.82, issue.3, pp.168-180, 2005.
DOI : 10.1016/j.aquabot.2005.04.003

D. S. Wilson, Introduction: Multilevel Selection Theory Comes of Age, The American Naturalist, vol.150, issue.S1, pp.1-4, 1997.
DOI : 10.1086/286046

C. C. Wu, P. K. Diggle, and W. E. Friedman, Kin recognition within a seed and the effect 636 of genetic relatedness of an endosperm to its compatriot embryo on maize seed 637 development, Proceedings of the National Academy of Sciences, pp.2217-2222, 2013.