M. Begon, J. Harper, and J. Townsend, Ecology: Individuals, populations and communities, p.1068, 1996.

C. Darwin, On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life, 1859.

E. Pianka, Sympatry of Desert Lizards (Ctenotus) in Western Australia, Ecology, vol.50, issue.6, 1969.
DOI : 10.2307/1936893

F. Ishtiaq, S. Javed, M. Coulter, and A. Rahmani, Resource Partitioning in Three Sympatric Species of Storks in Keoladeo National Park, India, Waterbirds, vol.33, issue.1, pp.41-49, 2010.
DOI : 10.1675/063.033.0105

J. Croxall and P. Prince, Food, feeding ecology and ecological segregation of seabirds at South Georgia, Biological Journal of the Linnean Society, vol.14, issue.1, 1980.
DOI : 10.1111/j.1095-8312.1980.tb00101.x

T. Schoener, Resource Partitioning in Ecological Communities, Science, vol.185, issue.4145, pp.27-39, 1974.
DOI : 10.1126/science.185.4145.27

G. Hunt and D. Schneider, Scale-dependent processes in the physical and biological environment of marine birds In: Seabirds, feeding ecology and role in marine ecosystems, pp.7-41, 1987.

A. Gaston, Seabirds: A natural history, 2004.

G. Orians and N. Pearson, On the theory of central place foraging In: DJ HornRD MitchellGR Stairs. Analysis of ecological systems, pp.155-177, 1979.

S. Lewis, T. Sherratt, K. Hamer, and S. Wanless, Evidence of intraspecific competition for food in a pelagic seabird, Nature, vol.412, 2001.

H. Young, D. Mccauley, R. Dirzo, R. Dunbar, and S. Shaffer, Niche partitioning among and within sympatric tropical seabirds revealed by stable isotope analysis, Marine Ecology Progress Series, vol.416, pp.285-294, 2010.
DOI : 10.3354/meps08756

E. Frere, F. Quintana, P. Gandini, and R. Wilson, Foraging behaviour and habitat partitioning of two sympatric cormorants in Patagonia, Argentina, Ibis, vol.9, issue.3, pp.558-564, 2008.
DOI : 10.1111/j.1474-919X.2008.00824.x

J. Navarro, M. Forero, J. González-solís, J. Igual, and J. Bécares, Foraging segregation between two closely related shearwaters breeding in sympatry, Biology Letters, vol.128, issue.2, 2009.
DOI : 10.1016/j.ympev.2006.05.006

H. Weimerskirch, S. Shaffer, Y. Tremblay, D. Costa, and H. Gadenne, Species- and sex-specific differences in foraging behaviour and foraging zones in blue-footed and brown boobies in the Gulf of California, Marine Ecology Progress Series, vol.391, pp.267-278, 2009.
DOI : 10.3354/meps07981

URL : https://hal.archives-ouvertes.fr/hal-00511439

M. Forero, J. González-solís, K. Hobson, J. Donazar, and M. Bertellotti, Stable isotopes reveal trophic segregation by sex and age in the southern giant petrel in two different food webs, Marine Ecology Progress Series, vol.296, pp.107-113, 2005.
DOI : 10.3354/meps296107

S. Bearhop, R. Phillips, M. Mcgill, Y. Cherel, and D. Dawson, Stable isotopes indicate sex-specific and long-term individual foraging specialisation in diving seabirds, Marine Ecology Progress Series, vol.311, pp.157-164, 2006.
DOI : 10.3354/meps311157

URL : https://hal.archives-ouvertes.fr/hal-00182601

T. Guilford, R. Wynn, M. Mcminn, A. Rodríguez, and A. Fayet, Geolocators Reveal Migration and Pre-Breeding Behaviour of the Critically Endangered Balearic Shearwater Puffinus mauretanicus, PLoS ONE, vol.13, issue.2, p.22470471, 2012.
DOI : 10.1371/journal.pone.0033753.g005

A. Moody and K. Hobson, Alcid winter diet in the northwest Atlantic determined by stable isotope analysis, Mar Ornithol, vol.35, pp.39-46, 2007.

J. González-solís, J. Croxall, and V. Afanasyev, Offshore spatial segregation in giant petrelsMacronectes spp.: differences between species, sexes and seasons, Aquatic Conservation: Marine and Freshwater Ecosystems, vol.6, issue.S1, pp.22-36, 2008.
DOI : 10.1002/aqc.911

Y. Cherel, K. Hobson, C. Guinet, and C. Vanpe, Stable isotopes document seasonal changes in trophic niches and winter foraging individual specialization in diving predators from the Southern Ocean, Journal of Animal Ecology, vol.75, issue.44, pp.826-836, 2007.
DOI : 10.1007/s00442-003-1270-z

URL : https://hal.archives-ouvertes.fr/hal-00185936

J. Thiebot, Y. Cherel, P. Trathan, and C. Bost, Coexistence of oceanic predators on wintering areas explained by population-scale foraging segregation in space or time, Ecology, vol.93, issue.1, 2012.
DOI : 10.1034/j.1600-048X.2000.310302.x

URL : https://hal.archives-ouvertes.fr/hal-00624598

J. Thiebot, Y. Cherel, P. Trathan, and C. Bost, Inter-population segregation in the wintering areas of macaroni penguins, Marine Ecology Progress Series, vol.421, pp.279-290, 2011.
DOI : 10.3354/meps08907

URL : https://hal.archives-ouvertes.fr/hal-00558229

J. Roscales, E. Gomez-diaz, V. Neves, and J. González-solís, Trophic versus geographic structure in stable isotope signatures of pelagic seabirds breeding in the northeast Atlantic, Marine Ecology Progress Series, vol.434, pp.1-13, 2011.
DOI : 10.3354/meps09211

D. Boertmann, A. Mosbech, K. Falk, and K. Kampp, eabird colonies in western Greenland (60 o -79 o 30'N. lat.). National Environmental Research Institute: Denmark, 1996.

K. Kampp and K. Falk, The birds of Ydre Kitsissut (Kitsissut Avaliit), Southwest Greenland. Meddelelser om Grønland, BioScience, vol.42, pp.1-25, 1994.

J. Linnebjerg, Foraging ecology of breeding seabirds in Greenland, 2012.

R. Wilson, J. Ducamp, G. Rees, B. Culik, and K. Niekamp, Estimation of location: global coverage using light intensity Wildlife telemetry: remote monitoring and tracking of animals, pp.131-134, 1992.

R. Hill, Theory of geolocation by light levels Elephant seals: population ecology, behaviour and physiology, pp.227-236, 1994.

P. Ekstrom, An advance in geolocation by light. Memoirs of the National Institute of Polar Research, pp.210-226, 2004.

M. Frederiksen, B. Moe, F. Daunt, R. Phillips, and R. Barrett, Multicolony tracking reveals the winter distribution of a pelagic seabird on an ocean basin scale, Diversity and Distributions, vol.163, issue.6, 2012.
DOI : 10.1111/j.1472-4642.2011.00864.x

R. Phillips, J. Silk, J. Croxall, V. Afanasyev, and D. Briggs, Accuracy of geolocation estimates for flying seabirds, Marine Ecology Progress Series, vol.266, pp.265-272, 2004.
DOI : 10.3354/meps266265

K. Hamer, E. Humphreys, S. Garthe, J. Hennicke, and G. Peters, Annual variation in diets, feeding locations and foraging behaviour of gannets in the North Sea: flexibility, consistency and constraint, Marine Ecology Progress Series, vol.338, pp.295-305, 2007.
DOI : 10.3354/meps338295

B. Worton, Kernel Methods for Estimating the Utilization Distribution in Home-Range Studies, Ecology, vol.70, issue.1, pp.164-168, 1989.
DOI : 10.2307/1938423

K. Elliott, G. Davoren, and A. Gaston, The influence of buoyancy and drag on the dive behaviour of an Arctic seabird, the Thick-billed Murre, Canadian Journal of Zoology, vol.85, issue.3, pp.352-36110, 2007.
DOI : 10.1139/Z07-012

C. Thaxter, S. Wanless, F. Daunt, M. Harris, and S. Benvenuti, Influence of wing loading on the trade-off between pursuit-diving and flight in common guillemots and razorbills, Journal of Experimental Biology, vol.213, issue.7, pp.1018-1025, 2010.
DOI : 10.1242/jeb.037390

D. Nettleship and T. Birkhead, The Atlantic Alcidae: The evolution, distribution and biology of the Auks inhabiting the Atlantic Ocean and adjacent water areas, 1985.

K. Hobson and R. Clark, Assessing Avian Diets Using Stable Isotopes II: Factors Influencing Diet-Tissue Fractionation, The Condor, vol.94, issue.1, pp.189-19710, 1992.
DOI : 10.2307/1368808

S. Carleton and R. Cmd, The effect of cold-induced increased metabolic rate on the rate of 13C and 15N incorporation in house sparrows (Passer domesticus), Oecologia, vol.124, issue.2, pp.226-232, 2005.
DOI : 10.1007/s00442-005-0066-8

S. Bearhop, S. Waldron, S. Votier, and R. Furness, Factors That Influence Assimilation Rates and Fractionation of Nitrogen and Carbon Stable Isotopes in Avian Blood and Feathers, Physiological and Biochemical Zoology, vol.75, issue.5, pp.451-45810, 2002.
DOI : 10.1086/342800

J. Kelly, Stable isotopes of carbon and nitrogen in the study of avian and mammalian trophic ecology, Canadian Journal of Zoology, vol.78, issue.1, pp.1-27, 2000.
DOI : 10.1139/z99-165

A. Gaston, P. Smith, L. Tranquilla, W. Montevecchi, and D. Fifield, Movements and wintering areas of breeding age Thick-billed Murre Uria lomvia from two colonies in Nunavut, Canada, Marine Biology, vol.116, issue.9, pp.1929-1941, 2011.
DOI : 10.1007/s00227-011-1704-9

W. Montevecchi, A. Hedd, M. Tranquilla, L. Fifield, D. Burke et al., Tracking seabirds to identify ecologically important and high risk marine areas in the western North Atlantic, Biological Conservation, vol.156, 2012.
DOI : 10.1016/j.biocon.2011.12.001

S. Wanless, M. Harris, and J. Morris, A Comparison of Feeding Areas Used by Individual Common Murres (Uria aalge), Razorbills (Alca torda) and an Atlantic Puffin (Fratercula arctica) during the Breeding Season, Colonial Waterbirds, vol.13, issue.1, pp.16-2410, 1990.
DOI : 10.2307/1521416

M. Bradstreet and R. Brown, Feeding ecology of the Atlantic Alcidae In: DN NettleshipTR Birkhead. The Atlantic Alcidae: The evolution, distribution and biology of the auks inhabiting the Atlantic Ocean and adjacent water areas, pp.264-318, 1985.

D. Thompson, K. Lilliendahl, J. Solmundsson, R. Furness, and S. Waldron, Trophic Relationships among Six Species of Icelandic Seabirds as Determined through Stable Isotope Analysis, The Condor, vol.101, issue.4, pp.898-903, 1999.
DOI : 10.2307/1370085

J. Lavers, M. Hipfner, G. Chapdelaine, and J. Hipfner, Razorbill (Alca torda) In: A Poole. The Birds of North America Online, 2009.

C. Barger and A. Kitaysky, Isotopic segregation between sympatric seabird species increases with nutritional stress, Biology Letters, vol.162, issue.1, pp.442-445, 2012.
DOI : 10.1007/s00442-007-0717-z

R. Barrett, M. Asheim, and V. Bakken, , breeding in the Barents Sea, Canadian Journal of Zoology, vol.75, issue.4, pp.618-63110, 1997.
DOI : 10.1139/z97-077

T. Birkhead and D. Nettleship, , at the Gannet Islands, Labrador. I. Morphometrics and timing of breeding, Canadian Journal of Zoology, vol.65, issue.7, pp.1621-162910, 1987.
DOI : 10.1139/z87-251

Y. Cherel, R. Phillips, K. Hobson, and R. Mcgill, Stable isotope evidence of diverse species-specific and individual wintering strategies in seabirds, Biology Letters, vol.74, issue.1400, p.17148388, 2006.
DOI : 10.1098/rspb.1998.0390

URL : https://hal.archives-ouvertes.fr/hal-00184875

K. Lilliendahl, Winter diets of auks in Icelandic coastal waters, Marine Biology Research, vol.13, issue.2, 2009.
DOI : 10.1006/jmsc.1997.0264

R. Elliot, P. Ryan, and W. Lidster, The winter diet of thick-billed murres in coastal Newfoundland waters, Stud Avian Biol, vol.14, pp.125-138, 1990.

B. Graham, P. Koch, S. Newsome, K. Mcmahon, and D. Aurioles, Using isoscapes to trace the movements and foraging behaviour of top predators in oceanic ecosystems Isoscapes: Understanding movement, pattern, and proscess on Earth through isotope mapping, pp.299-318, 2010.

A. Mosbech, F. Merkel, D. Boertmann, K. Falk, and M. Frederiksen, Thick-billed Murre studies in Disko Bay (Ritenbenk), 2009.

T. Clarke, A. Diamond, and J. Chardine, ) Wintering in the Outer Bay of Fundy Confirmed by Radio-Tracking, Waterbirds, vol.33, issue.4, pp.541-545, 2010.
DOI : 10.1675/063.033.0414

F. Huettmann, A. Diamond, B. Dalzell, and K. Macintosh, Winter distribution, ecology and movements of Razorbills Alca torda and other auks in the outer Bay of Fundy, Eastern Canada, Mar Ornithol, vol.33, pp.161-171, 2005.

T. Guilford, R. Freeman, D. Boyle, B. Dean, and H. Kirk, A Dispersive Migration in the Atlantic Puffin and Its Implications for Migratory Navigation, PLoS ONE, vol.278, issue.1713, p.21799734, 2011.
DOI : 10.1371/journal.pone.0021336.s027

D. Boertmann, P. Lyngs, F. Merkel, and A. Mosbech, The significance of Southwest Greenland as winter quarters for seabirds, Bird Conservation International, vol.14, issue.02, pp.87-112, 2004.
DOI : 10.1017/S0959270904000127

F. Merkel, A. Mosbech, D. Boertmann, and L. Grøndahl, Winter seabird distribution and abundance off south-western Greenland, 1999, Polar Research, vol.19, issue.1, pp.17-36, 1999.
DOI : 10.1111/j.1751-8369.2002.tb00064.x

URL : http://doi.org/10.3402/polar.v21i1.6471

P. Lyngs, Migration and winter ranges of birds in Greenland -An analysis of ringing recoveries, Dansk Ornitol Forenings Tidsskr, vol.97, pp.1-167, 2003.

C. Wernham, M. Toms, J. Marchant, J. Clark, and G. Siriwardena, The Migration Atlas -Movements of the birds of Britain and Ireland, p.883, 2002.

V. Bakken, O. Runde, and E. Tjørve, Norsk ringmerkingsatlas, 2003.

S. Lewis, D. Grémillet, F. Daunt, P. Ryan, and R. Crawford, Using behavioural and state variables to identify proximate causes of population change in a seabird, Oecologia, vol.75, issue.4, pp.606-614, 2006.
DOI : 10.1007/s00442-005-0321-z

URL : https://hal.archives-ouvertes.fr/hal-00110887