K. Amano and G. J. Vermeij, Taxonomy and evolution of the genus Ocinebrellus 379 (Gastropoda: Muricidae) in Japan, Paleontological Research, vol.2, pp.199-212, 1998.

H. Bandelt and P. Forster, Median-joining networks for inferring intraspecific phylogenies, Molecular Biology and Evolution, vol.16, issue.1, pp.37-48, 1999.
DOI : 10.1093/oxfordjournals.molbev.a026036

R. Barrett and P. D. Hebert, Identifying spiders through DNA barcodes, Canadian Journal of Zoology, vol.83, issue.3, 2005.
DOI : 10.1139/z05-024

V. Becquet and B. Simon-bouhet, Glacial refugium versus range limit: Conservation genetics of Macoma Balthica, a key species in the Bay of Biscay (France), Journal of Experimental Marine Biology and Ecology, vol.432, issue.433, p.386, 2012.
DOI : 10.1016/j.jembe.2012.07.008

URL : https://hal.archives-ouvertes.fr/hal-00872186

J. Bergsten, D. T. Bilton, and T. Fujisawa, The Effect of Geographical Scale of Sampling on DNA Barcoding, Systematic Biology, vol.61, issue.5, pp.851-869, 2012.
DOI : 10.1093/sysbio/sys037

V. Berrou and D. Merle, Comparative morphology of Pliocene, p.390, 2004.
URL : https://hal.archives-ouvertes.fr/hal-00326214

P. Bouchet and R. Houart, Ocenebra inornata (Récluz, 1851) Accessed through: World 394 Register of Marine Species at 395 http://www.marinespecies.org/aphia.php?, pp.403745-2014, 2014.

J. T. Carlton, Man's Role in Changing the Face of the Ocean: Biological Invasions and Implications for Conservation of Near-Shore Environments, Conservation Biology, vol.41, issue.3, pp.265-394, 1989.
DOI : 10.1016/0272-7714(88)90021-2

K. Amano and G. J. Vermeij, Taxonomy and evolution of the genus Ocinebrellus 395 (Gastropoda: Muricidae) in Japan, Paleontological Research, vol.2, pp.199-212, 1998.

H. Bandelt and P. Forster, Median-joining networks for inferring intraspecific phylogenies, Molecular Biology and Evolution, vol.16, issue.1, pp.37-48, 1999.
DOI : 10.1093/oxfordjournals.molbev.a026036

R. Barrett and P. D. Hebert, Identifying spiders through DNA barcodes, Canadian Journal of Zoology, vol.83, issue.3, 2005.
DOI : 10.1139/z05-024

V. Becquet and B. Simon-bouhet, Glacial refugium versus range limit: Conservation genetics of Macoma Balthica, a key species in the Bay of Biscay (France), Journal of Experimental Marine Biology and Ecology, vol.432, issue.433, p.402, 2012.
DOI : 10.1016/j.jembe.2012.07.008

URL : https://hal.archives-ouvertes.fr/hal-00872186

J. Bergsten, D. T. Bilton, and T. Fujisawa, The Effect of Geographical Scale of Sampling on DNA Barcoding, Systematic Biology, vol.61, issue.5, pp.851-869, 2012.
DOI : 10.1093/sysbio/sys037

V. Berrou and D. Merle, Comparative morphology of Pliocene, p.406, 2004.
URL : https://hal.archives-ouvertes.fr/hal-00326214

P. Bouchet and R. Houart, Ocenebra inornata (Récluz, 1851) Accessed through: World 410 Register of Marine Species at 411 http://www.marinespecies.org/aphia.php?, pp.403745-2014, 2014.

J. T. Carlton, Man's Role in Changing the Face of the Ocean: Biological Invasions and Implications for Conservation of Near-Shore Environments, Conservation Biology, vol.41, issue.3, pp.265-414, 1989.
DOI : 10.1016/0272-7714(88)90021-2

J. T. Carlton, Introduced marine and estuarine mollusks of North America: an end-of- 416 the-20th-century perspective, Journal of Shellfish Research, vol.11, pp.489-505, 1992.

X. De-montaudouin and P. Sauriau, Contribution to a synopsis of marine species 421 richness in the Pertuis Charentais Sea with new insights in soft-bottom macrofauna of 422, 2000.

A. Gómez and D. H. Lunt, Refugia within Refugia: Patterns of Phylogeographic 450, 2007.

A. Graham, Molluscs: Prosobranchs and Pyramidellid Gastropods: Keys and Notes 453 for the Identification of the Species, p.454, 1998.

M. G. Harasewych and S. L. Adamkewicz, NEOGASTROPOD PHYLOGENY: A MOLECULAR PERSPECTIVE, Journal of Molluscan Studies, vol.63, issue.3, pp.327-351, 1997.
DOI : 10.1093/mollus/63.3.327

P. D. Hebert and A. Cywinska, Biological identifications through DNA barcodes, Proceedings of the Royal Society B: Biological Sciences, vol.270, issue.1512, p.457, 2003.
DOI : 10.1098/rspb.2002.2218

. Bowen, Transcriptomic analysis of growth heterosis in larval Pacific oysters 460 (Crassostrea gigas), Proceedings of the National Academy of Sciences of the United States 461 of America, pp.2313-2318, 2007.

G. W. Hewitt, Some genetic consequences of ice ages, and their role in divergence and speciation, Biological Journal of the Linnean Society, vol.58, issue.3, pp.247-276, 1996.
DOI : 10.1111/j.1095-8312.1996.tb01434.x

G. M. Hewitt, Genetic consequences of climatic oscillations in the Quaternary, Philosophical Transactions of the Royal Society B: Biological Sciences, vol.359, issue.1442, p.465, 2004.
DOI : 10.1098/rstb.2003.1388

M. G. Hoskin, Effects of the East Australian Current on the genetic structure of a direct developing muricid snail (Bedeva hanleyi, Angas): variability within and among local populations, Biological Journal of the Linnean Society, vol.69, issue.2, pp.245-262, 2000.
DOI : 10.1111/j.1095-8312.2000.tb01201.x

J. Lützen and M. Faasse, The Japanese oyster drill Ocinebrellus inornatus (Récluz, Mollusca, Gastropoda, Muricidae), introduced to the Limfjord, pp.1851-497, 2012.

R. N. Mack and D. Simberloff, BIOTIC INVASIONS: CAUSES, EPIDEMIOLOGY, GLOBAL CONSEQUENCES, AND CONTROL, Ecological Applications, vol.10, issue.3, pp.690-710, 2000.
DOI : 10.1146/annurev.py.25.090187.001231

C. A. Maggs and R. Castilho, Evaluating signatures of glacial refugia for North 501, 2008.

C. Martel, Invasions biologiques et perturbations anthropiques des écosystèmes 503 littoraux : source, profil d'expansion et impacts d'un gastéropode marin, Ocinebrellus 504 inornatus, introduit accidentellement sur les côtes atlantiques françaises, p.505, 2003.

C. Martel and D. Bourguet, Invasion by the marine gastropod Ocinebrellus 507 inornatus in France. II. Expansion along the Atlantic coast, Marine Ecology Progress, vol.508, 2004.

C. Martel and J. M. Guarini, Invasion by the marine gastropod Ocinebrellus 510 inornatus in France. III. Comparison of biological traits with the resident species 511, 2004.

C. Martel and F. Viard, Invasion by the marine gastropod Ocinebrellus inornatus 513, 2004.

I. France, Scenario for the source of introduction, Journal of Experimental Marine, vol.514

J. B. Mitton and M. C. Grant, Associations Among Protein Heterozygosity, Growth Rate, and Developmental Homeostasis, Annual Review of Ecology and Systematics, vol.15, issue.1, pp.479-499, 1984.
DOI : 10.1146/annurev.es.15.110184.002403

H. A. Mooney and E. E. Cleland, The evolutionary impact of invasive species, Proceedings 518 of the National Academy of Sciences of the United States of America, pp.5446-5451, 2001.
DOI : 10.1073/pnas.091093398

M. Nei, Molecular Evolutionary Genetics, p.520, 1987.

E. Pante and B. , Simon-Bouhet (2013) marmap: A Package for Importing, Plotting and Analyzing 524

C. Parmesan and G. Yohe, A globally coherent fingerprint of climate change impacts across natural systems, Nature, vol.4, issue.6918, pp.37-42, 2003.
DOI : 10.1038/34073

P. Pascal, Rôle des espèces autochtones dans les processus d'invasions biologiques 528 marines: l', 2004.

. Oléron, Comptes-rendus de l'Académie des Sciences de Paris, pp.697-703

S. Ratnasingham and P. D. Hebert, BOLD : The Barcode of Life Data System BOLD, p.535, 2007.

F. Riquet and C. Daguin-thiébaut, Contrasting patterns of genome-wide 538, 2013.

G. M. Ruiz and T. K. Rawlings, Global spread of microorganisms by ships: ballast 541 water discharged from vessels harbours a cocktail of potential pathogens, Nature, vol.408, pp.542-591, 2000.

A. K. Sakai and F. W. Allendorf, The Population Biology of Invasive Species, Annual 544 rReview of Ecology and Systematics, pp.305-332, 2001.
DOI : 10.1146/annurev.ecolsys.32.081501.114037

H. Seebens and M. T. Gastner, The risk of marine bioinvasion caused by global 549 shipping, Ecology Letters, vol.6, pp.782-790, 2013.

B. Simon-bouhet and P. Garcia-meunier, Multiple introductions promote range expansion of the mollusc Cyclope neritea (Nassariidae) in France: evidence from mitochondrial sequence data, Molecular Ecology, vol.15, issue.6, pp.1699-1711, 2006.
DOI : 10.1111/j.1365-294X.2006.02881.x

URL : https://hal.archives-ouvertes.fr/hal-00485683

S. Y. Strauss and J. A. Lau, Evolutionary responses of natives to introduced species: 554 what do introductions tell us about natural communities? Ecology Letters, pp.357-374, 2006.

F. Tajima, Statistical method for testing the neutral mutation hypothesis by DNA 556 polymorphism, Genetics, vol.123, issue.3, pp.585-595, 1989.

J. D. Thompson and T. J. Gibson, The ClustalX windows interface: flexible strategies 558 for multiple sequence alignment aided by quality analysis tools, Nucleic Acid Research, vol.559, pp.24-4876, 1997.

M. J. Wittmann and M. Hutzenthaler, Ecological and genetic effects of introduced species on their native competitors, Theoretical Population Biology, vol.84, pp.25-35, 2013.
DOI : 10.1016/j.tpb.2012.11.003

S. M. Woolley and D. Posada, A Comparison of Phylogenetic Network Methods Using Computer Simulation, PLoS ONE, vol.132, issue.4, p.1913, 2008.
DOI : 10.1371/journal.pone.0001913.s001

S. Wright, The genetical structure of populations, Annals of Human Genetics, vol.15, pp.323-565, 1951.

S. Zou, Q. Li, and L. Kong, Multigene barcoding and phylogeny of geographically 570 widespread muricids (Gastropoda: Neogastropoda) along the coast of china, 2012.

E. Zouros, Associative overdominance: Evaluating the effects of inbreeding and linkage disequilibrium, Genetica, vol.42, issue.1-3, pp.35-46, 1993.
DOI : 10.1007/BF02424504