K. Alef, Nitrogen mineralization in soils, Methods in applied soil microbiology and biochemistry. Academic, pp.234-245, 1995.

M. Aubert, F. Bureau, D. Alard, and J. Bardat, Effect of tree mixture on the humic epipedon and vegetation diversity in managed beech forests (Normandy, France), Canadian Journal of Forest Research, vol.34, issue.1, pp.233-248, 2004.
DOI : 10.1139/x03-205

V. Bailey, J. Smith, and J. Bolton, Fungal-to-bacterial ratios in soils investigated for enhanced C sequestration, Soil Biology and Biochemistry, vol.34, issue.7, pp.997-1007, 2002.
DOI : 10.1016/S0038-0717(02)00033-0

J. Bauhus, D. Paré, and L. Coté, Effects of tree species, stand age and soil type on soil microbial biomass and its activity in a southern boreal forest, Soil Biology and Biochemistry, vol.30, issue.8-9, pp.1077-1089, 1997.
DOI : 10.1016/S0038-0717(97)00213-7

N. Bernier and J. Ponge, Humus form dynamics during the sylvogenetic cycle in a mountain spruce forest, Soil Biology and Biochemistry, vol.26, issue.2, pp.183-220, 1994.
DOI : 10.1016/0038-0717(94)90161-9
URL : https://hal.archives-ouvertes.fr/hal-00506004

M. Booth, J. Stark, and E. Rastetter, CONTROLS ON NITROGEN CYCLING IN TERRESTRIAL ECOSYSTEMS: A SYNTHETIC ANALYSIS OF LITERATURE DATA, Ecological Monographs, vol.75, issue.2, pp.139-157, 2005.
DOI : 10.1007/s003740050542

P. Bottner, F. Austrui, J. Cortez, G. Billes, and M. Couteaux, Decomposition of 14C- and 15N-labelled plant material, under controlled conditions, in coniferous forest soils from a north???south climatic sequence in western Europe, Soil Biology and Biochemistry, vol.30, issue.5, pp.597-610, 1998.
DOI : 10.1016/S0038-0717(97)00166-1

S. Brais, C. Camiré, Y. Bergeron, and D. Paré, Changes in nutrient availability and forest floor characteristics in relation to stand age and forest composition in the southern part of the boreal forest of northwestern Quebec, Forest Ecology and Management, vol.76, issue.1-3, pp.181-189, 1995.
DOI : 10.1016/0378-1127(95)03541-H

E. Brierley and M. Wood, Heterotrophic nitrification in an acid forest soil: isolation and characterisation of a nitrifying bacterium, Soil Biology and Biochemistry, vol.33, issue.10, pp.1403-1409, 2001.
DOI : 10.1016/S0038-0717(01)00045-1

L. Coté, S. Brown, D. Paré, J. Fyles, and J. Bauhus, Dynamics of carbon and nitrogen mineralization in relation to stand type, stand age and soil texture in the boreal mixedwood, Soil Biology and Biochemistry, vol.32, issue.8-9, pp.1079-1090, 2000.
DOI : 10.1016/S0038-0717(00)00017-1

D. Boer, W. Kowalchuk, and G. , Nitrification in acid soils: micro-organisms and mechanisms, Soil Biology and Biochemistry, vol.33, issue.7-8, pp.853-866, 2001.
DOI : 10.1016/S0038-0717(00)00247-9

L. Durin, J. Géhu, A. Noirfalise, and N. Sougnez, Les hêtraies atlantiques et leur essaim climatique dans le nord-ouest et l'ouest de la France, pp.66-89, 1967.

G. Gea-izquierdo, B. Allen-diaz, and A. Miguel, Canellas I (2010) How do trees affect spatio-temporal heterogeneity of nutrient cycling in mediterranean annual grasslands?, Ann For Sci, vol.67, issue.1, p.11

J. Gobat, M. Aragno, and W. Matthey, The living soil: fundamentals of soil science and soil biology, Science Publishers, U.S, p.602, 2004.

S. Hart, J. Stark, E. Davidson, and M. Firestone, Nitrogen mineralization, immobilization, and nitrification Methods of soil analysis part II: microbiological and biochemical Properties, pp.985-1017, 1994.

M. Hayatsu, K. Tago, and M. Saito, Various players in the nitrogen cycle: Diversity and functions of the microorganisms involved in nitrification and denitrification, Soil Science and Plant Nutrition, vol.61, issue.1, pp.33-45, 2008.
DOI : 10.1007/s002030000231

R. Hynes and R. Knowles, Commentary on ???Inhibition by acetylene of ammonia oxidation in Nitrosomonas europaea???, FEMS Microbiology Letters, vol.4, pp.319-321, 1978.
DOI : 10.1093/femsle/fnx068

Y. Inagaki, S. Miura, and A. Kohzu, Effects of forest type and stand age on litterfall quality and soil N dynamics in Shikoku district, southern Japan, Forest Ecology and Management, vol.202, issue.1-3, pp.107-117, 2004.
DOI : 10.1016/j.foreco.2004.07.029

B. Jabiol, A. Brêthes, J. Ponge, F. Toutain, and J. Brun, L'humus sous toutes ses formes, École Nationale du Génie Rural, des Eaux et Forêts, 2007.
URL : https://hal.archives-ouvertes.fr/hal-00508410

J. Jussy, M. Colin-belgrand, and J. Ranger, Production and root uptake of mineral nitrogen in a chronosequence of Douglas-fir (Pseudotsuga menziesii) in the Beaujolais Mounts, Forest Ecology and Management, vol.128, issue.3, pp.197-209, 2000.
DOI : 10.1016/S0378-1127(99)00147-4

A. Kooijman, J. Van-mourik, and M. Schilder, The relationship between N mineralization or microbial biomass N with micromorphological properties in beech forest soils with different texture and pH, Biology and Fertility of Soils, vol.8, issue.5, pp.449-459, 2009.
DOI : 10.1007/s00374-009-0354-2

B. Laignel, F. Quesnel, M. Lecoustumier, and R. Meyer, Variability of the clay fraction of the clay with flints of the western part of the Paris Basin, CR Acad Sci Paris, vol.326, pp.467-472, 1998.

E. Lang and G. Jagnow, Fungi of a forest soil nitrifying at low pH values, FEMS Microbiology Letters, vol.38, issue.5, pp.257-265, 1986.
DOI : 10.1111/j.1574-6968.1986.tb01736.x

P. Lavelle and A. Spain, Soil ecology, 2001.
DOI : 10.1007/978-94-017-5279-4
URL : https://hal.archives-ouvertes.fr/bioemco-00455666

X. Li, Y. Hu, S. Han, Y. Liu, and Y. Zhang, La chute de liti??re et la chimie de la liti??re changent au fil du temps dans une vieille for??t temp??r??e du Nord-Est de la Chine, Annals of Forest Science, vol.26, issue.2, p.8, 2010.
DOI : 10.1051/forest/2009099

J. Park, K. Kalbitz, and E. Matzner, Resource control on the production of dissolved organic carbon and nitrogen in a deciduous forest floor, Soil Biology and Biochemistry, vol.34, issue.6, pp.813-822, 2002.
DOI : 10.1016/S0038-0717(02)00011-1

H. Pedersen, K. Dunkin, and M. Firestone, The relative importance of autotrophic and heterotrophic nitrification in a conifer forest soil as measured by N-15 tracer and pool dilution techniques, Biogeochemistry, vol.44, issue.2, pp.135-150, 1999.

Z. Piotrowska-seget, R. Engel, E. Nowak, and J. Kozdroj, Successive soil treatment with captan or oxytetracycline affects non-target microorganisms, World Journal of Microbiology and Biotechnology, vol.1, issue.12, pp.2843-2848, 2008.
DOI : 10.1007/s11274-008-9815-2

J. Ponge, Humus forms in terrestrial ecosystems: a framework to biodiversity, Soil Biology and Biochemistry, vol.35, issue.7, pp.935-945, 2003.
DOI : 10.1016/S0038-0717(03)00149-4
URL : https://hal.archives-ouvertes.fr/hal-00498465

J. Ponge and L. Delhaye, The heterogeneity of humus profiles and earthworm communities in a virgin beech forest, Biology and Fertility of Soils, vol.93, issue.1, pp.24-32, 1995.
DOI : 10.1007/BF00307837
URL : https://hal.archives-ouvertes.fr/hal-00505505

R. Development and C. Team, R: a language and environment for statistical computing, R Foundation for Statistical Computing, 2008.

J. Schimel and J. Bennett, NITROGEN MINERALIZATION: CHALLENGES OF A CHANGING PARADIGM, Ecology, vol.85, issue.3, pp.591-602, 2004.
DOI : 10.1007/BF01814901

C. Ste-marie and D. Paré, Soil, pH and N availability effects on net nitrification in the forest floors of a range of boreal forest stands, Soil Biology and Biochemistry, vol.31, issue.11, pp.311579-1589, 1999.
DOI : 10.1016/S0038-0717(99)00086-3

J. Trap, F. Bureau, M. Akpa-vinceslas, C. R. , and A. M. , Changes in soil N mineralization and nitrification pathways along a mixed forest chronosequence, Forest Ecology and Management, vol.258, issue.7, pp.1284-1293, 2009.
DOI : 10.1016/j.foreco.2009.06.021

S. Welke and G. Hope, Influences of stand composition and age on forest floor processes and chemistry in pure and mixed stands of Douglas-fir and paper birch in interior British Columbia, Forest Ecology and Management, vol.219, issue.1, pp.29-42, 2005.
DOI : 10.1016/j.foreco.2005.08.040

D. Zaykin, L. Zhivotovsky, P. Westfall, and B. Weir, Truncated product method for combiningP-values, Genetic Epidemiology, vol.96, issue.2, pp.170-185, 2002.
DOI : 10.1002/gepi.0042

B. Zeller, S. Recous, M. Kunze, J. Moukoumi, M. Colin-belgrand et al., Influence of tree species on gross and net N transformations in forest soils, Annals of Forest Science, vol.64, issue.2, pp.151-158, 2007.
DOI : 10.1051/forest:2006099
URL : https://hal.archives-ouvertes.fr/hal-00884067

X. Zhao, Q. Lin, and P. Brookes, Does soil ergosterol concentration provide a reliable estimate of soil fungal biomass?, Soil Biology and Biochemistry, vol.37, issue.2, pp.311-317, 2005.
DOI : 10.1016/j.soilbio.2004.07.041