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F. , G. Laboratoire-de-microbiologie, G. Et-ecologie, and . Marines, F-13288 Marseille Cedex 9, France E-mail address: franck.gilbert@univmed.fr D. Cossa IFREMER, Centre de Nantes, BP 21105, F.44311 Nantes Cedex 03, France E-mail address: daniel.cossa@ifremer.fr F. Prévot Lab. Géochimie des Eaux, Institut de Physique du Globe de Paris, CNRS (UMR 7154)