C. Barnes, S. Jennings, and J. T. Barry, Environmental correlates of large-scale spatial variation in the ??13C of marine animals, Estuarine, Coastal and Shelf Science, vol.81, issue.3, pp.368-374, 2009.
DOI : 10.1016/j.ecss.2008.11.011

E. Bellier, B. Planque, and P. Petitgas, Historical fluctuations in spawning location of anchovy (Engraulis encrasicolus) and sardine (Sardina pilchardus) in the Bay of Biscay during 1967?73 and 2000?2004, Fisheries Oceanography, vol.33, issue.2, pp.1-15, 2000.
DOI : 10.1046/j.1365-2419.2003.00222.x

G. Cabana and J. B. Rasmussen, Comparison of aquatic food chains using nitrogen isotopes., Proceedings of the National Academy of Science, pp.10844-10847, 1996.
DOI : 10.1073/pnas.93.20.10844

S. Caut, E. Angulo, and F. Courchamp, C): the effect of diet isotopic values and applications for diet reconstruction, Journal of Applied Ecology, vol.75, issue.2, pp.443-453, 2009.
DOI : 10.1111/j.1365-2664.2009.01620.x

E. Chassot, T. Rouyer, V. M. Trenkel, and D. Gascuel, Investigating trophic-level variability in Celtic Sea fish predators, Journal of Fish Biology, vol.157, issue.4, pp.763-781, 2008.
DOI : 10.1111/j.1095-8649.2008.01938.x

URL : https://hal.archives-ouvertes.fr/ird-00548995

Y. Cherel, C. Fontaine, G. D. Jackson, C. H. Jackson, and P. Richard, Tissue, ontogenic and sex-related differences in ??13C and ??15N values of the oceanic squid Todarodes filippovae (Cephalopoda: Ommastrephidae), Marine Biology, vol.201, issue.4, pp.699-708, 2009.
DOI : 10.1007/s00227-008-1121-x

T. Chouvelon, J. Spitz, Y. Cherel, F. Caurant, R. Sirmel et al., Inter-specific and ontogenic differences in ??13C and ??15N values and Hg and Cd concentrations in cephalopods, Marine Ecology Progress Series, vol.433, pp.107-120, 2011.
DOI : 10.3354/meps09159

URL : https://hal.archives-ouvertes.fr/hal-00613955

P. M. Cury, L. J. Shannon, J. P. Roux, G. M. Daskalov, A. Jarre et al., Trophodynamic indicators for an ecosystem approach to fisheries, ICES Journal of Marine Science, vol.62, issue.3, pp.430-442, 2005.
DOI : 10.1016/j.icesjms.2004.12.006

L. A. Dehn, E. H. Follmann, D. L. Thomas, G. G. Sheffield, C. Rosa et al., Trophic relationships in an Arctic food web and implications for trace metal transfer, Science of The Total Environment, vol.362, issue.1-3, pp.103-123, 2006.
DOI : 10.1016/j.scitotenv.2005.11.012

D. Niro, M. J. Epstein, and S. , Mechanism of carbon fractionation associated with lipid synthesis, Science, vol.197, pp.261-263, 1977.

D. Niro, M. J. Epstein, and S. , Influence of diet on the distribution of carbon isotopes in animals, Geochimica et Cosmochimica Acta, vol.42, pp.495-506, 1978.

D. Niro, M. J. Epstein, and S. , Influence of diet on the distribution of nitrogen isotopes in animals, Geochimica et Cosmochimica Acta, vol.45, pp.341-351, 1981.

S. Dubois, F. Orvain, M. Leal, J. C. Ropert, M. Lefebvre et al., Small-scale spatial variability of food partitioning between cultivated oysters and associated suspension-feeding species, as revealed by stable isotopes, Marine Ecology Progress Series, vol.336, pp.151-160, 2007.
DOI : 10.3354/meps336151

URL : https://hal.archives-ouvertes.fr/hal-00481694

R. L. France, Carbon-13 enrichment in benthic compared to planktonic algae:foodweb implications, Marine Ecology Progress Series, vol.124, pp.307-312, 1995.
DOI : 10.3354/meps124307

B. Fry, Food web structure on Georges Bank from stable C, N, and S isotopic compositions, Limnology and Oceanography, vol.33, issue.5, pp.1182-1190, 1988.
DOI : 10.4319/lo.1988.33.5.1182

L. Z. Gannes, D. M. O-'brien, and C. Martínez-del-rio, STABLE ISOTOPES IN ANIMAL ECOLOGY: ASSUMPTIONS, CAVEATS, AND A CALL FOR MORE LABORATORY EXPERIMENTS, Ecology, vol.78, issue.4, pp.1271-1276, 1997.
DOI : 10.2307/278984

S. M. Garcia, A. Zerbi, T. Do-chi, and G. Lasserre, The ecosystem approach to fisheries. Issues, terminology, principles, institutional foundations, implementation and outlook, 2003.

D. Gascuel, Y. M. Bozec, E. Chassot, A. Colomb, and M. Laurans, The trophic spectrum: theory and application as an ecosystem indicator, ICES Journal of Marine Science, vol.62, issue.3, pp.443-452, 2005.
DOI : 10.1016/j.icesjms.2004.12.013

M. M. Guzzo, G. D. Haffner, S. Sorge, S. A. Rush, and A. T. Fisk, Spatial and temporal variabilities of ??13C and ??15N within lower trophic levels of a large lake: implications for estimating trophic relationships of consumers, Hydrobiologia, vol.646, issue.1, pp.41-53, 2011.
DOI : 10.1007/s10750-011-0794-1

S. Hansson, J. E. Hobbie, R. Elmgren, U. Larsson, B. Fry et al., THE STABLE NITROGEN ISOTOPE RATIO AS A MARKER OF FOOD-WEB INTERACTIONS AND FISH MIGRATION, Ecology, vol.78, issue.7, pp.2249-2257, 1997.
DOI : 10.2307/1941739

A. Herbland, D. Delmas, P. Laborde, B. Sautour, and F. Artigas, Phytoplankton spring bloom of the Gironde plume waters in the Bay of Biscay: early phosphorus limitation and food-web consequences, Oceanologica Acta, vol.21, issue.2, pp.279-291, 1998.
DOI : 10.1016/S0399-1784(98)80015-7

J. M. Hill, C. D. Mcquaid, and S. Kaehler, Biogeographic and nearshore??offshore trends in isotope ratios of intertidal mussels and their food sources around the coast of southern Africa, Marine Ecology Progress Series, vol.318, pp.63-73, 2006.
DOI : 10.3354/meps318063

J. Hjelm, L. Persson, and B. Christensen, Growth, morphological variation and ontogenetic niche shifts in perch (Perca fluviatilis) in relation to resource availability, Oecologia, vol.122, issue.2, pp.190-199, 2000.
DOI : 10.1007/PL00008846

K. A. Hobson, Tracing origins and migration of wildlife using stable isotopes: a review, Oecologia, vol.120, issue.3, pp.314-326, 1999.
DOI : 10.1007/s004420050865

K. A. Hobson and H. E. Welch, Determination of trophic relationships within a high Arctic marine food web using ??13C and ??15N analysis, Marine Ecology Progress Series, vol.84, pp.9-18, 1992.
DOI : 10.3354/meps084009

K. A. Hobson, A. Fisk, N. Karnovsky, M. Holst, J. M. Gagnon et al., A stable isotope (? 13 C, ? 15 N) model for the North Water food web: implications for evaluating trophodynamics and the flow of energy and contaminants. Deep-Sea Research Part II, pp.5131-5150, 2002.

S. Jennings, J. K. Pinnegar, N. V. Polunin, and T. V. Boon, Weak cross-species relationships between body size and trophic level belie powerful size-based trophic structuring in fish communities, Journal of Animal Ecology, vol.401, issue.Suppl. B, pp.934-944, 2001.
DOI : 10.1046/j.0021-8790.2001.00552.x

S. Jennings and K. J. Warr, Environmental correlates of large-scale spatial variation in the ??15N of marine animals, Marine Biology, vol.25, issue.6, pp.1131-1140, 2003.
DOI : 10.1046/j.1365-2435.1999.00301.x

V. S. Karpouzi and K. I. Stergiou, The relationships between mouth size and shape and body length for 18 species of marine fishes and their trophic implications, Journal of Fish Biology, vol.15, issue.6, pp.1353-1365, 2003.
DOI : 10.1046/j.1095-8649.2003.00118.x

J. F. Kelly, Stable isotopes of carbon and nitrogen in the study of avian and mammalian trophic ecology, Canadian Journal of Zoology, vol.78, issue.1, pp.1-27, 2000.
DOI : 10.1139/z99-165

C. Koutsikopoulos and B. Le-cann, Physical processes and hydrological structures related to the Bay of Biscay anchovy, Scientia Marina, vol.60, pp.9-19, 1996.

C. M. Kurle, E. H. Sinclair, A. E. Edwards, and C. J. Gudmundson, Temporal and spatial variation in the ??15N and ??13C values of fish and squid from Alaskan waters, Marine Biology, vol.82, issue.3, pp.2389-2404, 2011.
DOI : 10.1007/s00227-011-1741-4

S. Lefebvre, C. Harma, and J. L. Blin, Trophic typology of coastal ecosystems based on ??13C and ??15N ratios in an opportunistic suspension feeder, Marine Ecology Progress Series, vol.390, pp.27-37, 2009.
DOI : 10.3354/meps08187

URL : https://hal.archives-ouvertes.fr/hal-00480681

S. Lefebvre, M. Leal, J. C. Dubois, S. Orvain, F. Blin et al., Seasonal dynamics of trophic relationships among co-occurring suspension-feeders in two shellfish culture dominated ecosystems, Estuarine, Coastal and Shelf Science, vol.82, issue.3, pp.415-425, 2009.
DOI : 10.1016/j.ecss.2009.02.002

L. Loc-'h, F. Hily, C. Grall, and J. , Benthic community and food web structure on the continental shelf of the Bay of Biscay (North Eastern Atlantic) revealed by stable isotopes analysis, Journal of Marine Systems, vol.72, pp.17-34, 2008.

V. Lesage, M. O. Hammill, and K. M. Kovacs, Marine mammals and the community structure of the Estuary and Gulf of St Lawrence, Canada: evidence from stable isotope analysis, Marine Ecology Progress Series, vol.210, pp.203-221, 2001.
DOI : 10.3354/meps210203

P. Lorance, J. A. Bertrand, A. Brind-'amour, M. J. Rochet, and V. M. Trenkel, Assessment of impacts from human activities on ecosystem components in the Bay of Biscay in the early 1990s, Aquatic Living Resources, vol.22, issue.4, pp.409-431, 2009.
DOI : 10.1051/alr/2009049

P. Lorance, D. Latrouite, and B. Séret, Observations of Chondrichthyan fishes (sharks, rays and chimaeras) in the Bay of Biscay (North-Eastern Atlantic) from submersibles, Proceedings of the 3 rd European Elasmobranch Association Meeting, pp.29-45, 2000.

M. Lunven, J. F. Guillaud, A. Youénou, M. P. Crassous, R. Berric et al., Nutrient and phytoplankton distribution in the Loire River plume (Bay of Biscay, France) resolved by a new Fine Scale Sampler, Estuarine, Coastal and Shelf Science, vol.65, issue.1-2, pp.94-108, 2005.
DOI : 10.1016/j.ecss.2005.06.001

C. Martínez-del-rio, N. Wolf, S. A. Carleton, and L. Z. Gannes, Isotopic ecology ten years after a call for more laboratory experiments, Biological Reviews, vol.61, issue.1, pp.91-111, 2009.
DOI : 10.1111/j.1469-185X.2008.00064.x

J. M. Mcclelland, I. Valiela, and R. H. Michener, Nitrogen-stable isotope signatures in estuarine food webs: A record of increasing urbanization in coastal watersheds, Limnology and Oceanography, vol.42, issue.5, pp.930-937, 1997.
DOI : 10.4319/lo.1997.42.5.0930

J. H. Mccutchan, . Jr, W. M. Lewis, . Jr, C. Kendall et al., Variation in trophic shift for stable isotope ratios of carbon, nitrogen, and sulfur, Oikos, vol.102, issue.2, pp.378-390, 2003.
DOI : 10.1034/j.1600-0706.2003.12098.x

P. Mèndez-fernandez, P. Bustamante, A. Bode, T. Chouvelon, M. Ferreira et al., Foraging ecology of five toothed whale species in the Northwest Iberian Peninsula, inferred using carbon and nitrogen isotope ratios, Journal of Experimental Marine Biology and Ecology, vol.413, pp.150-158, 2012.
DOI : 10.1016/j.jembe.2011.12.007

F. Ménard, A. Lorrain, M. Potier, and F. Marsac, Isotopic evidence of distinct feeding ecologies and movement patterns in two migratory predators (yellowfin tuna and swordfish) of the western Indian Ocean, Marine Biology, vol.79, issue.2, pp.141-152, 2007.
DOI : 10.1007/s00227-007-0789-7

L. Meynier, C. Pusineri, J. Spitz, M. B. Santos, G. J. Pierce et al., Intraspecific dietary variation in the short-beaked common dolphin Delphinus delphis in the Bay of Biscay: importance of fat fish, Marine Ecology Progress Series, vol.354, pp.277-287, 2008.
DOI : 10.3354/meps07246

R. H. Michener and L. Kaufman, Stable isotope ratios as tracers in marine food webs: an update Stable isotopes in ecology and environmental science, pp.238-282, 2007.

M. Minagawa and E. Wada, Stepwise enrichment of 15N along food chains: Further evidence and the relation between ??15N and animal age, Geochimica et Cosmochimica Acta, vol.48, issue.5, pp.1135-1140, 1984.
DOI : 10.1016/0016-7037(84)90204-7

J. B. Mion, R. A. Stein, and E. A. Marschall, RIVER DISCHARGE DRIVES SURVIVAL OF LARVAL WALLEYE, Ecological Applications, vol.8, issue.1, pp.88-103, 1998.
DOI : 10.1146/annurev.es.15.110184.002141

J. P. Montoya, Natural abundance of 15 N in marine planktonic ecosystems Stable isotopes in ecology and environmental science, pp.176-201, 2007.

S. D. Newsome, C. Martinez-del-rio, S. Bearhop, and D. L. Phillips, A niche for isotopic ecology, Frontiers in Ecology and the Environment, vol.130, issue.8, pp.429-436, 2007.
DOI : 10.1126/science.1059412

L. Nøttestad, J. Giske, J. C. Holst, and G. Huse, A length-based hypothesis for feeding migrations in pelagic fish, Canadian Journal of Fisheries and Aquatic Sciences, vol.56, issue.S1, pp.26-34, 1999.
DOI : 10.1139/f99-222

M. L. Palomares and D. Pauly, SeaLifeBase, World Wide Web electronic publication, www.sealifebase.org, version, 2010.

D. Pauly, V. Christensen, J. Dalsgaard, R. Froese, F. C. Torres et al., Fishing Down Marine Food Webs, Science, vol.279, issue.5352, pp.860-863, 1998.
DOI : 10.1126/science.279.5352.860

D. Pauly and M. L. Palomares, Fishing down marine food web: it is far more pervasive than we thought, Bulletin of Marine Science, vol.76, pp.197-211, 2005.

D. Pauly and R. Watson, Background and interpretation of the 'Marine Trophic Index' as a measure of biodiversity, Philosophical Transactions of the Royal Society B: Biological Sciences, vol.5, issue.6863, pp.415-423, 2005.
DOI : 10.1038/35107050

B. J. Peterson and B. Fry, Stable Isotopes in Ecosystem Studies, Annual Review of Ecology and Systematics, vol.18, issue.1, pp.293-320, 1987.
DOI : 10.1146/annurev.es.18.110187.001453

J. K. Pinnegar and N. V. Polunin, Differential fractionation of delta13C and delta15N among fish tissues: implications for the study of trophic interactions, Functional Ecology, vol.13, issue.2, pp.225-231, 1999.
DOI : 10.1139/cjfas-54-5-1142

B. Planque, P. Lazure, and A. M. Jégou, Detecting hydrological landscapes over the Bay of Biscay continental shelf in spring, Climate Research, vol.28, pp.41-52, 2004.
DOI : 10.3354/cr028041

G. A. Polis, D. Strong, and R. , Food Web Complexity and Community Dynamics, The American Naturalist, vol.147, issue.5, pp.813-846, 1996.
DOI : 10.1086/285880

D. M. Post, USING STABLE ISOTOPES TO ESTIMATE TROPHIC POSITION: MODELS, METHODS, AND ASSUMPTIONS, Ecology, vol.83, issue.3, pp.703-718, 2002.
DOI : 10.4319/lo.1994.39.5.1030

I. Puillat, P. Lazure, A. M. Jégou, L. Lampert, and P. I. Miller, Hydrographical variability on the French continental shelf in the Bay of Biscay, during the 1990s, Continental Shelf Research, vol.24, issue.10, pp.1143-1163, 2004.
DOI : 10.1016/j.csr.2004.02.008

I. Puillat, P. Lazure, A. M. Jégou, L. Lampert, and P. I. Miller, Mesoscale hydrological variability induced by northwesterly wind on the French continental shelf of the Bay of Biscay, Scientia Marina, vol.70, issue.S1, pp.15-26, 2006.
DOI : 10.3989/scimar.2006.70s115

J. C. Quéro, Guide des Poissons de l'Atlantique Européen, Les guides du naturaliste, Delachaux and Niestlé, 2003.

A. T. Revill, J. W. Young, and M. Lansdell, Stable isotopic evidence for trophic groupings and bio-regionalization of predators and their prey in oceanic waters off eastern Australia, Marine Biology, vol.79, issue.4, pp.1241-1253, 2009.
DOI : 10.1007/s00227-009-1166-5

D. M. Schell, B. A. Barnett, and K. A. Vinette, Carbon and nitrogen isotope ratios in zooplankton of the Bering, Chukchi and Beaufort seas, Marine Ecology Progress Series, vol.162, pp.11-23, 1998.
DOI : 10.3354/meps162011

J. M. Semmens, G. T. Pecl, B. M. Gillanders, C. M. Waluda, E. K. Shea et al., Approaches to resolving cephalopod movement and migration patterns, Reviews in Fish Biology and Fisheries, vol.71, issue.1???2, pp.401-423, 2007.
DOI : 10.1007/s11160-007-9048-8

G. D. Sherwood and G. A. Rose, Stable isotope analysis of some representative fish and invertebrates of the Newfoundland and Labrador continental shelf food web, Estuarine, Coastal and Shelf Science, vol.63, issue.4, pp.537-549, 2005.
DOI : 10.1016/j.ecss.2004.12.010

J. Spitz, E. Richard, L. Meynier, C. Pusineri, and V. Ridoux, Dietary plasticity of the oceanic striped dolphin, Stenella coeruleoalba, in the neritic waters of the Bay of Biscay, Journal of Sea Research, vol.55, issue.4, pp.309-320, 2006.
DOI : 10.1016/j.seares.2006.02.001

J. Spitz, Y. Rousseau, and V. Ridoux, Diet overlap between harbour porpoise and bottlenose dolphin: An argument in favour of interference competition for food?, Estuarine, Coastal and Shelf Science, vol.70, issue.1-2, pp.259-270, 2006.
DOI : 10.1016/j.ecss.2006.04.020

URL : https://hal.archives-ouvertes.fr/hal-00150198

J. Spitz, Y. Cherel, S. Bertin, J. Kiszka, A. Dewez et al., Prey preferences among the community of deep-diving odontocetes from the Bay of Biscay, Northeast Atlantic. Deep-Sea Res Part I 58, pp.273-282, 2011.
URL : https://hal.archives-ouvertes.fr/hal-00573185

V. M. Trenkel, O. Berthelé, P. Lorance, J. Bertrand, A. Brind-'amour et al., Atlas des grands invertébrés et poissons observés par les campagnes scientifiques, Ifremer, vol.100, pp.9-12, 2008.

A. Vanderklift and S. Ponsard, Sources of variation in consumer-diet ?15N enrichment: a meta-analysis, Oecologia, vol.136, issue.2, pp.169-182, 2003.
DOI : 10.1007/s00442-003-1270-z

M. J. Vander-zanden and W. W. Fetzer, Global patterns of aquatic food chain length, Oikos, vol.65, issue.8, pp.1378-1388, 2007.
DOI : 10.1111/j.0030-1299.2007.16036.x

M. J. Vander-zanden and J. B. Rasmussen, N AND THE TROPHIC POSITION OF AQUATIC CONSUMERS, Ecology, vol.80, issue.4, pp.1395-1404, 1999.
DOI : 10.2307/1941739

S. Vizzini and A. Mazzola, The effects of anthropogenic organic matter inputs on stable carbon and nitrogen isotopes in organisms from different trophic levels in a southern Mediterranean coastal area, Science of The Total Environment, vol.368, issue.2-3, pp.723-731, 2006.
DOI : 10.1016/j.scitotenv.2006.02.001