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G. Fichou and . Irmar, 35042 Rennes Cedex, France E-mail address: goulwen.fichou@univ-rennes1 Université de Bretagne Occidentale, 6, Av. Victor Le Gorgeu France E-mail address: Johannes.Huisman@univ-brest.fr Frédéric Mangolte d'Angers E-mail address: frederic.mangolte@univ-angers, fr Johannes Huisman, LMBA (UMR 6205), 29238.