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4 ? bourget@math.univ-angers.fr L. Chaumont 4 ? loic.chaumont@univ-angers.fr N. Sapoukhina 1,2,3 ? natalia.sapoukhina@angers.inra.fr 1 INRA, UMR1345 Institut de Recherche en Horticulture et Semences ? IRHS, SFR 4207, PRES UNAM, 42 rue Georges Morel, F-49071 Beaucouzé Cedex, France 2 AgroCampus-Ouest, UMR1345 Institut de Recherche en Horticulture et Semences ? IRHS, F-49045 ,